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Bolander's groundsel, Bolander's ragwort

plains butterweed, prairie groundsel

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Biennials or perennials, 20–60+ cm; rhizomatous and/or fibrous-rooted (bases erect to suberect), sometimes stoloniferous (mostly eastern populations).
Stems

1 or 2–3, loosely clustered, glabrous.

1 or 2–3, clustered, floccose-tomentose proximally and in leaf axils, otherwise sparsely tomentose or glabrescent.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(and proximal cauline) petiolate;

blades narrowly elliptic to elliptic-ovate or oblanceolate to suborbiculate or sublyrate, 20–70+ × 10–30+ mm, bases tapering to rounded or abruptly contracted, margins subentire to crenate, serrate-dentate, or pinnately lobed (abaxial faces floccose-tomentose, especially along midribs, ± glabrescent).

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually reduced (petiolate, sublyrate or pinnatisect, abaxial faces sparsely hairy; distals sessile, subentire to irregularly dissected).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

conspicuously bracteate, sparsely to densely tomentose.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

8–10;

corolla laminae 9–10 mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

60–70+;

corolla tubes 2.5–3.5 mm, limbs 3.5–4.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

13 or 21, green (tips sometimes cyanic), 5–6+ mm, densely tomentose proximally, glabrescent distally.

Calyculi

conspicuous.

inconspicuous.

Heads

8–15+ in corymbiform or subumbelliform arrays.

6–20+ in open or congested, corymbiform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

1.5–2.5 mm, usually hirtellous, sometimes glabrous;

pappi 6.5–7.5 mm.

2n

= 46.

= 46, 92.

Packera bolanderi

Packera plattensis

Phenology Flowering mid Apr–early Jun(–mid Jul, north).
Habitat Prairies, meadows, open wooded areas, along highways, railroads, around mining and construction areas, usually on limestone
Elevation 50–1800 m (200–5900 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CO; GA; IA; IL; IN; KS; LA; MI; MN; MO; MS; MT; ND; NE; NM; OH; OK; PA; SD; TN; VA; WI; WY; ON; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera plattensis is abundant, widespread, and almost weedy. Putative hybrids with other species are known. Plants in mesic, remnant prairies in the east are sometimes stoloniferous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 596.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio plattensis, Senecio pseudotomentosus
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Nuttall) W. A. Weber & Á. Löve: Phytologia 49: 48. (1981)
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