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Bolander's groundsel, Bolander's ragwort

northern groundsel

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials, 6–20+ cm; fibrous-rooted or rhizomatous (rhizomes horizontal to suberect, stout).
Stems

1 or 2–3, loosely clustered, glabrous.

usually 1, sometimes multiple, bases and leaf axils tomentose, otherwise glabrous.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(and proximal cauline) petiolate;

blades oblanceolate to obovate or lyrate (pinnately lobed to pinnatifid, terminal lobes larger than laterals), 10–30+ × 10–20+ mm, bases tapering, ultimate margins ± crenate to serrate.

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually to abruptly reduced (sessile; pinnatisect or entire).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

bracteate (bractlets purple-tinged), glabrous or glabrate.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

0 (rarely) or 10–12;

corolla laminae 8–10 mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

45–60+;

corolla tubes 3.5–4 mm, limbs 3–3.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

13, green (sometimes purple-tinged distally), 6–8 mm, glabrous.

Calyculi

conspicuous.

0 or inconspicuous (bractlets purple-tinged).

Heads

8–15+ in corymbiform or subumbelliform arrays.

1–2(–5+) in cymiform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

1.5–2.5 mm, hirtellous on ribs;

pappi 6–7 mm.

2n

= 46.

= 46.

Packera bolanderi

Packera hyperborealis

Phenology Flowering mid Jun–early Aug.
Habitat Open, dry, rocky or sandy areas, sometimes in fertile soils overlaying limestone
Elevation 0–900 m (0–3000 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; NT; NU; YT
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera hyperborealis is known only from lowland tundra from Alaska to the Mackenzie River and as far south as the northern end of the Franklin Mountains. It has also been collected as far north as Banks Island.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 589.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio hyperborealis
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Greenman) Á. Löve & D. Löve: Bot. Not. 128: 520. (1976)
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