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Bolander's groundsel, Bolander's ragwort

Hart's ragwort

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials, 40–70 cm; fibrous-rooted, sometimes stoloniferous (caudices ascending).
Stems

1 or 2–3, loosely clustered, glabrous.

1 or 2–3, clustered, bases and leaf axils usually loosely floccose-tomentose, sometimes glabrate.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

petiolate;

blades ovate to obovate or suborbiculate, 30–80 × 20–40+ mm, bases tapering, sometimes obtuse or contracted, margins serrate-dentate to weakly crenate (abaxial faces sometimes hairy).

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually reduced (petiolate, often sublyrate, sometimes weakly pinnatisect; mids and distals sessile, pinnatisect to subentire).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

bracteate, loosely tomentose or glabrescent.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

8 or 13;

corolla laminae 5–8 mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

50–65+;

corolla tubes 2–4 mm, limbs 2.5–3.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

(13–)21, green, 5–7 mm, glabrous or sparsely tomentose.

Calyculi

conspicuous.

0.

Heads

8–15+ in corymbiform or subumbelliform arrays.

3–9+ in open, corymbiform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

1–2 mm, glabrous;

pappi 3.5–4.5 mm.

2n

= 46.

= 44.

Packera bolanderi

Packera hartiana

Phenology Flowering mid May–early Jul.
Habitat Meadows or open areas, woodlands, along streams
Elevation 1600–2600 m [5200–8500 ft]
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera hartiana resembles both P. paupercula and P. plattensis; it is separated from both on the basis of geography and morphologic characters. New Mexico populations were formerly treated as P. quaerens; the morphologic characters used to separate the two entities vary and overlap.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 589.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio hartianus, P. quaerens, S. prionophyllus, S. quaerens
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (A. Heller) W. A. Weber & Á. Löve: Phytologia 49: 47. (1981)
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