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Bolander's groundsel, Bolander's ragwort

splitleaf groundsel

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials, 10–30+ cm; fibrous-rooted (caudices horizontal to erect, relatively short and stout).
Stems

1 or 2–3, loosely clustered, glabrous.

usually 1, rarely 2–3, clustered, glabrous.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(relatively thick and turgid) petiolate (petioles broad-winged or narrow, delicate);

blades ovate, ovate-orbiculate, oblong-lanceolate, orbiculate, suborbiculate, reniform, or subreniform, 10–50+ × 10–40 mm, bases tapering or subcordate to abruptly contracted, margins entire or crenate.

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually reduced (proximal and mid sometimes as large as or larger than basal, sessile, oblanceolate to obovate, sometimes lyrate, sometimes auriculate and/or clasping, subentire to irregularly dissected; distal usually sessile, bractlike, sometimes larger).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

inconspicuously bracteate or ebracteate, glabrous.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

8 or 13;

corolla laminae (deep yellow to orange) 5–8+ mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

45–60+;

corolla tubes 2–3 mm, limbs 2.5–3.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

(8–)13 or 21, green (tips cyanic), 5–7+ mm, glabrous.

Calyculi

conspicuous.

conspicuous (bases of bractlets swollen).

Heads

8–15+ in corymbiform or subumbelliform arrays.

1–6+ in congested, corymbiform or subumbelliform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

0.75–1.5 mm, glabrous, pappi 3–4 mm.

2n

= 46.

= 46.

Packera bolanderi

Packera dimorphophylla

Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; MT; NM; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
1. Basal leaves: petioles broad-winged, blades ovate to oblong-lanceolate, bases tapering; stems usually not branched
var. dimorphophylla
1. Basal leaves: petioles narrowly, if at all winged, blades suborbiculate, orbiculate, ovate-orbiculate, subreniform, or reniform, bases subcordate to abruptly contracted; stems branched or not branched
→ 2
2. Plants 30–50 cm; stems not branched; petiole lengths 1–3 times blades; Utah
var. intermedia
2. Plants 10–35 cm; stems frequently 1–3 times branched; petiole lengths 1–2 times blades; Idaho, Montana, Nevada, Wyoming
var. paysonii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 584.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
P. dimorphophylla var. dimorphophylla, P. dimorphophylla var. intermedia, P. dimorphophylla var. paysonii
Synonyms Senecio bolanderi Senecio dimorphophyllus
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Greene) W. A. Weber & Á. Löve: Phytologia 49: 46. (1981)
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