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Bolander's groundsel, Bolander's ragwort

dwarf arctic butterweed, high alpine butterweed, northwestern groundsel

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials, 4–10+ cm; rhizomatous and/or fibrous-rooted (mat-forming, bases ascending to erect, coarse).
Stems

1 or 2–3, loosely clustered, glabrous.

1 or 2–3, clustered, bases floccose-tomentose, leaf axils tomentose, glabrous elsewhere.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(thick, fleshy) petiolate;

blades usually ovate, oblong, or spatulate, sometimes sublyrate, 20–50+ × 20–40+ mm, bases tapering (to winged petioles) or abruptly contracted to subcordate (petioles narrow), margins crenate, coarsely serrate, or subentire.

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

(often cyanic) gradually reduced (sessile, not clasping; lanceolate to linear, usually irregularly and shallowly lobed, rarely entire).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

conspicuously bracteate, glabrous.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

10–12;

corolla laminae 8–14+ mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

55–75+;

corolla tubes 2.5–3.5 mm, limbs 3–4 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

21, deep red or green (tips reddish), 8–12+ mm, white-tomentose proximally.

Calyculi

conspicuous.

conspicuous (tips of bractlets often purple).

Heads

8–15+ in corymbiform or subumbelliform arrays.

1–2(–5+).

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

1–1.25 mm, glabrous;

pappi 4–7 mm.

2n

= 46.

= 160+.

Packera bolanderi

Packera contermina

Phenology Flowering early Jul–late Aug.
Habitat Subalpine or alpine, open areas, rocky slopes or ravines, moist tundra or snowbeds
Elevation 2100–3000 m (6900–9800 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
MT; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera contermina grows in rocky areas and produces relatively short rhizomes and abundant thin fibrous roots. In mesic sites, the rhizomes are more robust and the fibrous roots are fewer. This taxon has been treated as part of P. cymbalaria or P. subnuda. Morphologic and cytologic data lend support to its recognition at species rank.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 582.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio conterminus
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Greenman) J. F. Bain: Novon 9: 457. (1999)
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