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Bolander's groundsel, Bolander's ragwort

Brewer's ragwort

Habit Perennials, 20–50+ cm; fibrous-rooted (caudices creeping). Perennials or biennials, 40–100+ cm; fibrous-rooted (caudices erect, stout).
Stems

1 or 2–3, loosely clustered, glabrous.

1, glabrous or leaf axils tomentose.

Basal leaves

(and proximal cauline) petiolate;

blades ± orbiculate to oblong, or lyrate (lateral lobes 1–3 pairs, sinuses usually rounded, midribs usually winged), 50–120+ × 30–70+ mm, bases cordate to subcordate, margins shallowly to deeply incised.

(and proximal cauline) petiolate;

blades spatulate to obovate (pinnately lobed, terminal lobes ovate to oblong, lateral lobes 2–6+ pairs, smaller, midribs narrowly winged), 100–300+ × 20–50(–80+) mm, bases contracted to tapering, ultimate margins crenate or dentate to lacerate (faces glabrous).

Cauline leaves

gradually reduced (petiolate or sessile and clasping; lanceolate, incised).

gradually reduced (petiolate or sessile; obovate, irregularly incised to subpinnate, terminal lobes narrow).

Peduncles

ebracteate or inconspicuously bracteate, glabrous.

usually ebracteate (rarely with 1–2 bractlets), glabrous.

Ray florets

8 or 13;

corolla laminae 10–15+ mm.

8–10+;

corolla laminae 10–15+ mm.

Disc florets

30–45+;

corolla tubes 3–4 mm, limbs 3.5–4.5 mm.

45–60+;

corolla tubes 3–3.5 mm, limbs 4–4.5 mm.

Phyllaries

(8), 13, or 21, dark green, 5–8 mm, glabrous or sparsely tomentose proximally.

13 or 21, green, 7–9+ mm, glabrous.

Calyculi

conspicuous.

inconspicuous.

Heads

8–15+ in corymbiform or subumbelliform arrays.

15–50+ in corymbiform arrays.

Cypselae

2.5–3.5 mm, glabrous;

pappi 5–7 mm.

2–2.5 mm, glabrous;

pappi 6–7 mm.

2n

= 46.

= 46.

Packera bolanderi

Packera breweri

Phenology Flowering mid Apr–late May.
Habitat Dry, rocky soils, partially shaded or protected areas, grasslands, oak savannas, roadsides, disturbed areas
Elevation 200–1500 m (700–4900 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

The two varieties of Packera bolanderi have been treated as species. The morphologic characteristics used to distinguish the varieties vary widely within some populations. Presence or absence of multicellular hairs has been used to separate the varieties; that trait is also variable. The morphologic distinction is weak; habitat and elevation records lend support for recognizing two varieties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Packera breweri is known only from Coast Ranges from San Francisco Bay to Los Angeles County and along the eastern edges of the San Joaquin Valley. Populations are relatively small and sporadic; the plants appear not to hybridize with other species of Packera.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal leaves (relatively thick and turgid); phyllaries 13 or 21, 6–8+ mm, hairy proximally (hairs tan or brownish, multicelled)
var. bolanderi
1. Basal leaves (relatively thin); phyllaries 8, 13, (or 21), 5–6 mm, glabrous
var. harfordii
Source FNA vol. 20, p. 579. FNA vol. 20, p. 580.
Parent taxa Asteraceae > tribe Senecioneae > Packera Asteraceae > tribe Senecioneae > Packera
Sibling taxa
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. breweri, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
P. anonyma, P. antennariifolia, P. aurea, P. bernardina, P. bolanderi, P. cana, P. cardamine, P. castoreus, P. clevelandii, P. contermina, P. crocata, P. cymbalaria, P. cynthioides, P. debilis, P. dimorphophylla, P. eurycephala, P. fendleri, P. flettii, P. franciscana, P. ganderi, P. glabella, P. greenei, P. hartiana, P. hesperia, P. hyperborealis, P. indecora, P. ionophylla, P. layneae, P. macounii, P. malmstenii, P. millefolium, P. millelobata, P. multilobata, P. musiniensis, P. neomexicana, P. obovata, P. ogotorukensis, P. pauciflora, P. paupercula, P. plattensis, P. porteri, P. pseudaurea, P. quercetorum, P. sanguisorboides, P. schweinitziana, P. spellenbergii, P. streptanthifolia, P. subnuda, P. tampicana, P. texensis, P. tomentosa, P. tridenticulata, P. werneriifolia
Subordinate taxa
P. bolanderi var. bolanderi, P. bolanderi var. harfordii
Synonyms Senecio bolanderi Senecio breweri
Name authority (A. Gray) W. A. Weber & Á. Löve: Phytologia 49: 45. (1981) (Burtt Davy) W. A. Weber & Á. Löve: Phytologia 49: 46. (1981)
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