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ironwood, Knowlton's hop-hornbeam, western hop-hornbeam

Habit Trees, to 9 m; crowns open, narrowly rounded.
Bark

brownish gray, broken into narrow vertical scales or rather ragged strips.

thin, close or becoming furrowed or broken into plates;

lenticels not conspicuous.;

bark and wood tanniferous.

Twigs

sparsely to moderately pubescent, often with stipitate glands.

Leaves

blade ovate or broadly ovate-elliptic to broadly elliptic or nearly orbiculate, 2.5–6.5 × 1.5–5 cm, base narrowly rounded to cordate or cuneate, margins sharply and unevenly doubly serrate, apex acute;

surfaces abaxially pubescent, especially on veins.

2-ranked.

Inflorescences

staminate catkins 2–3 cm; pistillate catkins 0.6–1 cm.

Staminate flowers

perianth absent.

Pistillate flowers

2 per bract;

perianth adnate to ovary, often visible as membranaceous fringe at summit;

ovules with 2 integuments.

Infructescences

2.5–4 × 1.8–2.5 cm;

bracts 1–1.8 × 0.5–1 cm.

usually longer than 4 cm, consisting of relatively uncrowded clusters with large, nearly foliaceous bracts;

bracts deciduous with fruits.

Fruits

tiny to moderately large nuts, not winged;

pericarp thick and bony.

Trunk

and branches terete.

Young

twigs and buds usually without prominent, large, resinous glands;

pith circular to remotely triangular in cross section.

Ostrya knowltonii

Betulaceae subfam. coryloideae

Phenology Flowering late spring.
Habitat Streamsides and rocky slopes in moist canyons
Elevation 1200–2400 m (3900–7900 ft)
Distribution
from FNA
AZ; NM; TX; UT
[WildflowerSearch map]
[BONAP county map]
Primarily boreal and cool temperate zones of Northern Hemisphere
Discussion

Ostrya knowltonii occurs sporadically throughout the arid Southwest, including both rims of the Grand Canyon. On the basis of morphology and phytogeography, it appears to be more closely allied with Ostrya in mountainous western Mexico than with the eastern O. virginiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 4, species ca. 45 (3 genera, 7 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 3. FNA vol. 3.
Parent taxa Betulaceae > subfam. Coryloideae > Ostrya Betulaceae
Sibling taxa
O. chisosensis, O. virginiana
Subordinate taxa
Synonyms O. baileyi
Name authority Coville: Gard. & Forest 7: 114. (1894) Koehne: Deut. Dendrol., 106. (1893)
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