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orthotrichum moss, rock bristle-moss

orthotrichum moss

Habit Plants (1–)3–12.5 cm, light green to olive green. Plants 0.5–1 cm.
Stem

leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm;

margins recurved to narrowly revolute to near apex, entire;

apex sharply to slenderly acute;

basal laminal cells elongate to rectangular, walls thick, ± nodose;

distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low.

leaves stiff, erect-appressed or slightly flexuose when dry, lanceolate, 1.8–2.4 mm;

margins recurved to just below apex, entire;

apex acute;

basal laminal cells rectangular, walls thin, not nodose;

distal cells 8–12 µm, 1-stratose, smooth or papillae 1–3 per cell, conic, low.

Seta

to 1.8 mm.

0.7–1.5 mm.

Sexual condition

gonioautoicous.

gonioautoicous.

Capsule

immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length;

stomata superficial;

peristome single, rarely double;

prostome large, conspicuous, or rudimentary;

exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate;

endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened.

emergent, elliptic-cylindric and strangulate, 0.8–1.5 mm, strongly 8-ribbed, ribs thick, broad when dry, constricted below mouth when dry;

stomata immersed, 1/2 to completely covered by subsidiary cells, cells projecting, inner walls thickened;

peristome double;

prostome absent;

exostome teeth 8, reflexed to revolute when old and dry, finely and evenly papillose;

endostome segments 8 or 16, well developed, of 1 or 2 rows of cells, exterior surface smooth or finely striate.

Calyptra

oblong, smooth, hairy or rarely naked, hairs finely papillose.

oblong, smooth, rarely naked or hairs few, hairs variably smooth or papillose by projecting walls.

Spores

13–21 µm.

10–15 µm.

Specialized

asexual reproduction absent.

asexual reproduction by gemmae on abaxial surface of leaves.

Vaginula

with hairs present.

Orthotrichum rupestre

Orthotrichum stramineum

Habitat Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade Trees
Elevation low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) low to moderate elevations (10-500 m) (low to moderate elevations (0-1600 ft))
Distribution
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
[WildflowerSearch map]
from FNA
NF; Europe; Asia
Discussion

In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Orthotrichum stramineum is distinguished by its rather long, elliptic (fat), strongly ribbed capsules that are strangulate. The hairy vaginula and the shape of the capsule separate O. stramineum from O. stellatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 65. FNA vol. 28, p. 69.
Parent taxa Orthotrichaceae > Orthotrichum Orthotrichaceae > Orthotrichum
Sibling taxa
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
Synonyms Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum
Name authority Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) Hornschuch: in S. É. von Bridel, Bryol. Univ. 1: 789. (1827)
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