Orthotrichum rupestre |
Orthotrichum stellatum |
|
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orthotrichum moss, rock bristle-moss |
stellate orthotrichum moss |
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Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants 0.3–0.8 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves stiff, loosely erect-appressed when dry, lanceolate to oblong-lanceolate, 1.8–2.2 mm; margins recurved to just below apex, entire; apex narrowly obtuse to bluntly acute, not incurved when dry; basal laminal cells rectangular, walls thin, nodose toward costa; distal cells 8–13 µm, 1-stratose, lumina irregular due to uneven wall thickenings, smooth or papillae 1–3 per cell, conic, small. |
Seta | to 1.8 mm. |
0.4–1 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
emergent, oblong-cylindric and strangulate when dry, 0.4–1 mm, 8-ribbed when dry, ribs thick, broad, strongly constricted below mouth; stomata immersed, mid capsule, 1/2 to completely covered by subsidiary cells, cells projecting horizontally over stomata, inner walls thickened; peristome double; prostome absent; exostome teeth 8, reflexed, finely and evenly papillose; endostome segments 8, well developed, of 1 or 2 rows of cells, smooth. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
oblong, smooth, naked or sparsely hairy, hairs smooth. |
Spores | 13–21 µm. |
13–18 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae on abaxial surface of leaves. |
Vaginula | with hairs absent. |
|
Orthotrichum rupestre |
Orthotrichum stellatum |
|
Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Hardwood trunks in moist forests, with Orthotrichum ohioense in mesic, shaded stream edges |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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AR; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON
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Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum stellatum is distinguished by the strongly ribbed, strangulate capsules with well-developed endostomes of eight segments. In comparison, O. ohioense has weakly ribbed capsules that are not constricted below the mouth and an endostome that is lacking or rudimentary. From O. pumilum, the narrowly lanceolate leaves and small laminal cells with unevenly incrassate walls differentiate O. stellatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 68. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | O. braunii, O. strangulatum var. stellatum |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | Bridel: Bryol. Univ. 1: 274. (1826) |
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