Orthotrichum rupestre |
Orthotrichum spjutii |
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orthotrichum moss, rock bristle-moss |
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Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants to 3.5 cm, light green to olive green. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves stiff, loosely erect-appressed to erect-curved when dry, broadly lanceolate, ovate-lanceolate, or ovate-ligulate, 2–4 mm; margins strongly revolute from near base to just before apex, papillose-crenulate; apex bluntly acute to obtuse; basal laminal cells long-rectangular to elongate, walls thick, ± nodose; distal cells 10–15 µm, 1-stratose, papillae 1–3 per cell, 2-fid, large. |
Seta | to 1.8 mm. |
1–1.5 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
emergent, often shortly so, oblong to short-cylindric, 1.5–2 mm, nearly smooth to slightly 8-ribbed, sometimes deeply 8-ribbed and constricted with flaring mouth when old and dry; stomata superficial; peristome rudimentary, appearing absent; prostome absent; exostome teeth 16, erect, sometimes reflexed when old and dry, papillose; endostome segments occasionally rudimentary, usually absent. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
ovoid-campanulate, smooth, hairy, hairs papillose. |
Spores | 13–21 µm. |
8–11 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Orthotrichum rupestre |
Orthotrichum spjutii |
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Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Shaded forests near stream beds, canyons, mountains, deciduous trees, shaded rock |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | high elevations (2600 m) (high elevations (8500 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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CA |
Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum spjutii is best recognized by: large multifid distal laminal cell papillae, strongly revolute leaf margins, emergent, oblong capsules with peristome reduced to one or two cells (seemingly absent), and small spores. The marginal laminal cells are narrower than the basal cells and somewhat papillose, and the exostome teeth are very short and blunt. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 68. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | D. H. Norris & Vitt: Nova Hedwigia 56: 260, fig. 1. (1993) |
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