Orthotrichum rupestre |
Orthotrichum pusillum |
|
---|---|---|
orthotrichum moss, rock bristle-moss |
orthotrichum moss |
|
Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants 0.2–0.4 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves loosely appressed when dry, ovate-lanceolate to oblong-lanceolate, 1–2.2 mm; margins revolute nearly to apex, entire except for denticulate apex on some leaves; apex narrowly obtuse to broadly acute, sometimes apiculate; basal laminal cells broadly rectangular, walls thin, not nodose; distal cells 9–14 µm, 1-stratose, papillae 2–4 per cell, conic, small. |
Seta | to 1.8 mm. |
to 1 mm. |
Sexual condition | gonioautoicous. |
autoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
immersed to 1/2 emergent, oblong to ovate-oblong, 0.8–1.4 mm, smooth when dry, wrinkled to slightly 8-plicate when old, constricted below mouth when dry; stomata immersed; peristome single; prostome absent; exostome teeth 16, erect or recurved with age, finely papillose, rarely striate at apices; endostome segments usually absent, or 8, rudimentary, of 1 row of cells, smooth. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
oblong, smooth, naked. |
Spores | 13–21 µm. |
13–20 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae on abaxial surface of leaves. |
Orthotrichum rupestre |
Orthotrichum pusillum |
|
Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Trunks of deciduous trees in open, dry woods, roadsides, on Juniperus along streams |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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AL; AR; CT; DC; FL; GA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WV |
Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum pusillum is distinguished by its smooth, oblong capsules sharply contracted to the seta, 16 exostome teeth erect or recurved when old, some leaves of each plant having 1–4 small denticulations at the apex, and distal laminal cells 9–14 µm. The laminal cells are smaller than those of O. pumilum and in longitudinal rows, but they are larger, with thinner walls, than those of O. ohioense. Orthotrichum pusillum has a denticulate leaf apex that may be narrowly obtuse or acute, and laminal cells 9–14 µm with thin walls; O. pumilum has an apiculate leaf apex that is normally acute and entire, and laminal cells 14–20 µm with thin walls; and O. ohioense has an obtuse leaf apex that is never apiculate or denticulate, and laminal cells 8–10 µm with thick walls. Capsules of O. pusillum have stomata at mid capsule or beyond. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 64. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | O. psilocarpum |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | Mitten: J. Linn. Soc., Bot. 8: 25. (1864) |
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