Orthotrichum rupestre |
Orthotrichum pallens |
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orthotrichum moss, rock bristle-moss |
orthotrichum moss, pale bristle-moss |
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Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants 0.3–1 cm. | ||||||||
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves loosely erect-appressed and slightly incurved when dry, oblong-lanceolate to broadly lanceolate, 1.6–3 mm; margins recurved to near apex, entire or crenulate at apex; apex narrowly obtuse, acute, or apiculate; basal laminal cells rectangular, walls thin, not nodose; distal cells 9–14 µm, 1-stratose, papillae 2 or rarely 3 per cell, conic, low. |
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Seta | to 1.8 mm. |
0.5–1.5 mm. |
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Sexual condition | gonioautoicous. |
cladautoicous. |
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Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
emergent to barely exserted, oblong to oblong-cylindric, 1–2.5 mm, strongly 8-ribbed, narrowly cylindric and contracted along entire length when old; stomata immersed, to 1/2 covered by subsidiary cells, cells not projecting, inner walls not much thickened; peristome double; prostome absent; exostome teeth 8, erect, reflexed or recurved-reflexed with age, finely papillose, densely papillose, or papillose-striate; endostome segments 8 or 16, well developed, of 1 or 2 rows of cells, smooth or finely papillose. |
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Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
conic-oblong, smooth, naked or sparsely hairy, hairs smooth. |
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Spores | 13–21 µm. |
10–20 µm. |
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Specialized | asexual reproduction absent. |
asexual reproduction occasional, by gemmae on leaves. |
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Orthotrichum rupestre |
Orthotrichum pallens |
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Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | |||||||||
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | |||||||||
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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North America; Mexico; Europe |
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Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 9 (3 in the flora). Orthotrichum pallens is distinguished by the immersed stomata that are scarcely covered by thin-walled subsidiary cells. The peristome is double with a well-developed endostome of 16 (or eight) segments that remains intact until the capsule is quite old. Gametophytically, O. pallens is distinguished by narrowly obtuse, oblong-lanceolate leaves, and an autoicous sexual condition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 61. | ||||||||
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | O. immersum, O. pallens var. immersum | ||||||||
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | Bruch ex Bridel: Bryol. Univ. 1: 788. (1827) | ||||||||
Web links |