Orthotrichum rupestre |
Orthotrichum norrisii |
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orthotrichum moss, rock bristle-moss |
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Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants to 1.5 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves stiff, erect-appressed when dry, narrowly lanceolate to lanceolate-ligulate, 1.3–3 mm; margins recurved, becoming incurved or plane at apex, entire; apex narrowly obtuse, acute to notched and channeled; basal laminal cells rectangular, occasionally polygonal, walls thin to occasionally thick, rarely nodose; distal cells 10–16 µm, 1-stratose, papillae 2 or 3 per cell, simple, low. |
Seta | to 1.8 mm. |
0.5–1 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
emergent, cylindric when dry, 1.5–2 mm, slightly 8-ribbed entire length, ribs thin, delicate when dry, constricted below mouth when dry; stomata immersed, 1/2 to completely covered by subsidiary cells, cells projecting, inner walls thickened; peristome double; prostome absent; exostome teeth 8, reflexed, finely papillose; endostome segments 8, well developed, of 1 (or 2) rows of cells, smooth to faintly papillose. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
oblong-conic, smooth, hairy distally, hairs smooth. |
Spores | 13–21 µm. |
14–18 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction sometimes present by gemmae on leaves. |
Vaginula | with hairs present. |
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Orthotrichum rupestre |
Orthotrichum norrisii |
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Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Epiphytic in woodlands and chaparral |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | moderate elevations (200-900 m) (moderate elevations (700-3000 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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CA |
Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Orthotrichum norrisii is somewhat similar to O. tenellum, both species having a well-developed peristome of eight reflexed exostome teeth and eight long-lasting endostome segments. Also, both species occur on tree trunks in Mediterranean-like climates of California. Orthotrichum norrisii differs in having some leaves with irregular, channeled apices (versus mostly obtuse in O. tenellum) and slightly ribbed capsules (compared to thick, broad ribs in O. tenellum). The stomata are found in the proximal third of the capsule. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 59. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | F. Lara: Bryologist 111: 670, figs. 1 – 12. (2008) |
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