Orthotrichum rupestre |
Orthotrichum lyellii |
|
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orthotrichum moss, rock bristle-moss |
Lyell's bristle-moss, Lyell's orthotrichum moss |
|
Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants to 13 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves erect-spreading to loosely erect, ± contorted when dry, lanceolate to linear-lanceolate, 2.5–6.5 mm; margins plane, entire; apex narrowly acute to acuminate; basal laminal cells narrowly elongate, walls thick, ± nodose; distal cells 7–14 µm, 1-stratose, papillae 1–3 per cell, conic, large. |
Seta | to 1.8 mm. |
0.8–2.7 mm. |
Sexual condition | gonioautoicous. |
dioicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
immersed, emergent, or rarely exserted, oblong-cylindric, 1.7–3 mm, strongly 8-ribbed to mid capsule or entire length; stomata superficial; peristome double; prostome absent; exostome teeth 16, reflexed-recurved, acuminate, densely papillose; endostome segments 16, well developed, usually present when capsule is old and dry, thick, stout, of 2 rows of cells, narrower than exostome teeth, yellowish brown, coarsely papillose. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
conic-oblong, smooth, very hairy, hairs papillose. |
Spores | 13–21 µm. |
18–35 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae on leaves. |
Orthotrichum rupestre |
Orthotrichum lyellii |
|
Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Trees, covering trunks to 10 m, boulders |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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AK; CA; ID; OR; WA; BC; Mexico (Baja California); Europe; n Africa
|
Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum lyellii is easily distinguished from all other species of the genus by its dioicous sexual condition, the male plants being slightly smaller and having numerous and conspicuous perigonia. In addition, the 16, slenderly acuminate, whitish exostome teeth and 16 endostome segments are characteristic. Sterile plants usually have abundant laminal gemmae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 59. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | Dorcadion lyellii, O. lyellii var. howei, O. lyellii var. papillosum, O. lyellii var. pringlei, O. lyellii subsp. strictum, O. lyellioides, O. menziesii, O. papillosum, O. papillosum subsp. strictum, O. pringlei |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | Hooker & Taylor: Muscol. Brit., 76, plate 22 [upper left]. (1818) |
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