Orthotrichum rupestre |
Orthotrichum bartramii |
|
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orthotrichum moss, rock bristle-moss |
Bartram's orthotrichum moss |
|
Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants to 1 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves erect-appressed when dry, oblong to oblong-lanceolate, 2–3 mm; margins recurved to broadly revolute, incurved to form channel at apex, serrate at apex; apex cuspidate; basal laminal cells narrowly rectangular, walls thin, not nodose; distal cells 8–13 µm, 1-stratose, papillae 2 per cell, conic, small. |
Seta | to 1.8 mm. |
to 1 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous or cladautoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
± emergent or rarely immersed, oblong to oblong-cylindric when old, 1.3–2 mm, 8-ribbed entire length when dry, constricted below mouth when dry; stomata immersed; peristome double; prostome absent; exostome teeth 8, reflexed, densely and evenly papillose; endostome segments 8, not well developed, of 1 or 2 rows of cells, papillose, sometimes coarsely so. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
oblong-conic, smooth, hairy, hairs papillose. |
Spores | 13–21 µm. |
14–24 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Orthotrichum rupestre |
Orthotrichum bartramii |
|
Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Bark of Quercus or Populus tremuloides |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | high elevations (3000 m) (high elevations (9800 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
|
AZ; Mexico; Central America (Guatemala) |
Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The channeled, cuspidate apex of the mature leaves distinguishes Orthotrichum bartramii from all other species with immersed stomata. Stomata are found in the proximal capsule, half to completely covered by subsidiary cells. The sporophytic characters strongly resemble those of O. tenellum; however, the latter species has non-channeled, incurved leaf apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 52. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | R. S. Williams: Bryologist 28: 76, plate 9. (1925) |
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