Orthotrichum rupestre |
Orthotrichum affine |
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orthotrichum moss, rock bristle-moss |
orthotrichum moss, wood bristle-moss |
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Habit | Plants (1–)3–12.5 cm, light green to olive green. | Plants to 3 cm. |
Stem | leaves stiff, erect-appressed when dry, narrowly lanceolate to ovate-lanceolate, 2–4.5 mm; margins recurved to narrowly revolute to near apex, entire; apex sharply to slenderly acute; basal laminal cells elongate to rectangular, walls thick, ± nodose; distal cells 6–13 µm, 1-stratose, papillae 1 or 2 per cell, conic and small, or 2-fid and low. |
leaves erect-appressed and straight when dry, ligulate, elongate- to ovate-lanceolate, 2–4 mm; margins recurved-revolute to just below apex, entire; apex acute; basal laminal cells long-rectangular to elongate, walls thick, nodose; distal cells 9–13 µm, 1-stratose, papillae 2 or 3 per cell, 2-fid, low. |
Seta | to 1.8 mm. |
1–2.2 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | immersed, emergent, or slightly exserted, usually 1/3 emergent, globose ovate, ovate-oblong, or short-oblong, 1.3–1.8 mm, slightly or distinctly 8-ribbed 1/2–2/3 length; stomata superficial; peristome single, rarely double; prostome large, conspicuous, or rudimentary; exostome teeth 16, erect or sometimes spreading-recurved when old, smooth, scattered-papillose, or coarsely papillose-granulate; endostome segments absent, or rarely 8, rudimentary, of 1 row of cells, smooth or roughened. |
1/2 emergent to short-exserted, long-cylindric when mature, cylindric and constricted along entire length when old and dry, 1.6–2.5 mm, strongly 8-ribbed 2/3 capsule length; stomata superficial; peristome double; prostome absent; exostome teeth 8, erect when mature, reflexed to reflexed-recurved when old, coarsely and obscurely papillose or sometimes papillose-reticulate, deeply perforate, cancellate; endostome segments 8, rarely 16, well developed, usually present when capsule is old and dry, narrow, of 2 rows of cells, papillose. |
Calyptra | oblong, smooth, hairy or rarely naked, hairs finely papillose. |
conic-oblong, smooth, hairs few, papillose. |
Spores | 13–21 µm. |
15–18 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction absent. |
Orthotrichum rupestre |
Orthotrichum affine |
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Habitat | Non-calcareous boulders and cliff faces in mesic areas of pine, spruce-fir, or aspen forests, base of trees, subalpine shade | Deciduous trees, shaded rock, shaded forests near stream beds, canyons, mountains |
Elevation | low to high elevations (100-3000 m) (low to high elevations (300-9800 ft)) | low to high elevations (0-3000 m) (low to high elevations (0-9800 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NT; NU; ON; YT; South America; e Africa; Pacific Islands (New Zealand); Atlantic Islands (Canary Islands); Greenland; n Africa; e Asia (Japan); Europe; Australia
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CA; CO; ID; MT; OR; UT; WA; AB; BC; Europe; Asia; n Africa
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Discussion | In typical form, Orthotrichum rupestre is characterized as much-branched plants in loose tufts with lanceolate leaves, apex narrowly and bluntly acute, basal cells elongate, thick-walled, and nodose, yellowish in older leaves, capsule oblong, emergent, 8-ribbed to mid capsule when old and dry, smooth or slightly 8-ribbed when mature, exostome of 16 erect teeth, coarsely papillose, and calyptra with abundant, spinulose, papillose hairs, some of which extend over the beak. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum affine, a species confined to western North America in the flora area, is best recognized by: superficial stomata; cylindric, strongly ribbed, almost exserted capsules; a well-developed but not stout endostome; and a calyptra that is only sparsely hairy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 65. | FNA vol. 28, p. 50. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion rupestre, O. bullatum, O. californicum, O. douglasii, O. macfaddenae, O. rupestre var. globosum, O. rupestre var. macfaddenae, O. texanum, O. texanum var. globosum | |
Name authority | Schleicher ex Schwagrichen: Sp. Musc. Frond. Suppl. 1(2): 27, plate 53 [top]. (1816) | Schrader ex Bridel: Muscol. Recent. 2(2): 22. (1801) |
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