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blunt-leaf bristle-moss, obtuse-leaf aspen moss

Bolander's orthotrichum moss

Habit Plants 0.3–1.4 cm. Plants 0.5–4 cm, dark green to black.
Stem

leaves erect-appressed when dry, ovate to ovate-lanceolate, 1.4–2 mm;

margins erect-incurved, entire;

apex obtuse;

basal laminal cells ± rectangular, walls thick, not nodose;

distal cells 13–18(–20) µm, 1-stratose, papillae 1 per cell, conic, large.

leaves stiff, erect-appressed, rarely incurved when dry, lanceolate, 2–3.5 mm;

margins plane, entire;

apex narrowly obtuse to acute;

basal laminal cells elongate, walls thick, ± nodose;

distal cells 7–11 µm, 2-stratose, papillae 2–4 per cell, conic, sometimes large.

Seta

0.5 mm.

1 mm.

Sexual condition

dioicous.

gonioautoicous.

Capsule

1/2 emergent, oblong-cylindric, 1.6–2 mm, strongly 8-ribbed to base;

stomata superficial;

peristome double;

prostome absent;

exostome teeth 8, reflexed, evenly papillose, sometimes irregularly striate distally;

endostome segments 8, well developed, of 2 rows of cells, smooth.

emergent, ovate-oblong when mature, ovate-cylindric when old, 1.4–2 mm, slightly 8-ribbed in distal 1/3 of capsule;

stomata superficial;

peristome double;

prostome present, rudimentary;

exostome teeth 8, erect, sometimes reflexed when old and dry, papillose to coarsely papillose-striate;

endostome segments 8, occasionally rudimentary, of 2 rows of cells, papillose.

Calyptra

short-conic, papillose, naked, not plicate.

oblong-conic, smooth, hairy, hairs papillose.

Spores

20–25 µm.

17–23 µm.

Specialized

asexual reproduction by gemmae on both surfaces of leaf.

asexual reproduction absent.

Orthotrichum obtusifolium

Orthotrichum bolanderi

Habitat Trunks of Populus, Juniperus, aspen, base of scrub oaks, river bottoms Rock, especially sandstone
Elevation low to high elevations (10-2000 m) (low to high elevations (0-6600 ft)) low to high elevation (100-2000 m) (low to high elevation (300-6600 ft))
Distribution
from FNA
AK; AZ; CA; CO; IA; ID; MD; ME; MI; MN; MT; NC; ND; NE; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; ON; PE; QC; SK; YT; Europe; Asia
[WildflowerSearch map]
from FNA
CA; WA; Mexico; Central America (Guatemala)
Discussion

Orthotrichum obtusifolium is a pronounced xerophyte growing only in pioneer situations. This species might be confused with O. exiguum, O. flowersii, and O. gymnostomum, but these species do not have distal laminal cells with one papilla per cell as do those of O. obtusifolium. In addition, O. exiguum and O. flowersii have reflexed to revolute leaf margins, and both are autoicous. Orthotrichum gymnostomum has distinctly involute leaf margins and concave leaves and is dioicous. The capsule stomata of O. obtusifolium are found at or beyond mid capsule.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Orthotrichum bolanderi is the only species with superficial stomata that has 2-stratose distal laminal cells. The plants have leaves with subsheathing bases, ovoid-cylindric capsules with long wrinkled necks, and reflexed-recurved exostome teeth. When dry, the leaves are stiffly erect-appressed; they lie flat on a microscope slide when wet.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 59. FNA vol. 28, p. 52.
Parent taxa Orthotrichaceae > Orthotrichum Orthotrichaceae > Orthotrichum
Sibling taxa
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
Synonyms Nyholmiella obtusifolia, O. inflexum
Name authority Bridel: Muscol. Recent. 2(2): 23. (1801) Sullivant: Icon. Musc., suppl., 64, plate 46. (1874)
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