Orthotrichum lyellii |
Orthotrichum underwoodii |
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Lyell's bristle-moss, Lyell's orthotrichum moss |
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Habit | Plants to 13 cm. | Plants to 1 cm. |
Stem | leaves erect-spreading to loosely erect, ± contorted when dry, lanceolate to linear-lanceolate, 2.5–6.5 mm; margins plane, entire; apex narrowly acute to acuminate; basal laminal cells narrowly elongate, walls thick, ± nodose; distal cells 7–14 µm, 1-stratose, papillae 1–3 per cell, conic, large. |
leaves erect when dry, lanceolate, 2–3.2(–3.5) mm; margins recurved on one side in proximal 2/3, plane or slightly recurved in proximal 1/4 on other side, entire or papillose-crenulate; apex acute; basal laminal cells rectangular to almost quadrate, walls thin, usually not nodose; distal cells (7–)11–14(–16) µm, 1-stratose, papillae 2–4 per cell, simple to 2-fid, large. |
Seta | 0.8–2.7 mm. |
0.4–0.6 mm. |
Sexual condition | dioicous. |
gonioautoicous or cladautoicous. |
Capsule | immersed, emergent, or rarely exserted, oblong-cylindric, 1.7–3 mm, strongly 8-ribbed to mid capsule or entire length; stomata superficial; peristome double; prostome absent; exostome teeth 16, reflexed-recurved, acuminate, densely papillose; endostome segments 16, well developed, usually present when capsule is old and dry, thick, stout, of 2 rows of cells, narrower than exostome teeth, yellowish brown, coarsely papillose. |
2/3 emergent to short-exserted, cylindric, 2–2.3 mm, narrowly 8-ribbed entire length, constricted below mouth when dry; stomata immersed; peristome double; prostome present, rudimentary; exostome teeth 16, erect when mature, reflexed when old, irregularly papillose-striate; endostome segments 16, well developed, of 1 row of cells, filiform, papillose. |
Calyptra | conic-oblong, smooth, very hairy, hairs papillose. |
conic-oblong, smooth, sparsely hairy, hairs smooth. |
Spores | 18–35 µm. |
16–18(–19) µm. |
Specialized | asexual reproduction by gemmae on leaves. |
asexual reproduction absent. |
Orthotrichum lyellii |
Orthotrichum underwoodii |
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Habitat | Trees, covering trunks to 10 m, boulders | Trees |
Elevation | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) | elevation unknown |
Distribution |
AK; CA; ID; OR; WA; BC; Mexico (Baja California); Europe; n Africa
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CA |
Discussion | Orthotrichum lyellii is easily distinguished from all other species of the genus by its dioicous sexual condition, the male plants being slightly smaller and having numerous and conspicuous perigonia. In addition, the 16, slenderly acuminate, whitish exostome teeth and 16 endostome segments are characteristic. Sterile plants usually have abundant laminal gemmae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Orthotrichum underwoodii is known only from type material from the Santa Clara Mountains of Santa Clara County. The plants have slightly ribbed cylindric capsules with erect exostomes of 16 teeth and endostomes of 16 long and linear segments. In peristome features O. underwoodii is similar to O. diaphanum, but that species has hair-points on its leaves. Orthotrichum underwoodii could be confused with O. tenellum, but the latter species has much more strongly ribbed capsules, reflexed exostome, endostome of eight well-developed but short segments, and blunt, very stiff leaves with incurved apices. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 59. | FNA vol. 28, p. 71. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion lyellii, O. lyellii var. howei, O. lyellii var. papillosum, O. lyellii var. pringlei, O. lyellii subsp. strictum, O. lyellioides, O. menziesii, O. papillosum, O. papillosum subsp. strictum, O. pringlei | |
Name authority | Hooker & Taylor: Muscol. Brit., 76, plate 22 [upper left]. (1818) | F. Lara: Bryologist 104: 268, figs. 1 – 19. (2001) |
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