Orthotrichum lyellii |
Orthotrichum shawii |
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Lyell's bristle-moss, Lyell's orthotrichum moss |
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Habit | Plants to 13 cm. | Plants to 4 cm, light green to olive green. |
Stem | leaves erect-spreading to loosely erect, ± contorted when dry, lanceolate to linear-lanceolate, 2.5–6.5 mm; margins plane, entire; apex narrowly acute to acuminate; basal laminal cells narrowly elongate, walls thick, ± nodose; distal cells 7–14 µm, 1-stratose, papillae 1–3 per cell, conic, large. |
leaves stiff, erect-appressed when dry, lanceolate, 2.7–3.6 mm; margins broadly recurved, plane at apex, entire; apex short-acuminate to acute; basal laminal cells elongate, walls thick, nodose; distal cells 9–15 µm, 1-stratose, papillae 2–4 per cell, simple or 2-fid, large. |
Seta | 0.8–2.7 mm. |
to 0.4 mm. |
Sexual condition | dioicous. |
gonioautoicous. |
Capsule | immersed, emergent, or rarely exserted, oblong-cylindric, 1.7–3 mm, strongly 8-ribbed to mid capsule or entire length; stomata superficial; peristome double; prostome absent; exostome teeth 16, reflexed-recurved, acuminate, densely papillose; endostome segments 16, well developed, usually present when capsule is old and dry, thick, stout, of 2 rows of cells, narrower than exostome teeth, yellowish brown, coarsely papillose. |
immersed to emergent, ovate to cylindric-ovate, 1.6–1.8 mm, usually smooth or slightly 8-ribbed when dry; stomata superficial; peristome usually single; prostome small or absent; exostome teeth 16, recurved, coarsely papillose to papillose-striate; endostome rarely present, segments 8, rudimentary, of 1 row of cells, smooth. |
Calyptra | conic-oblong, smooth, very hairy, hairs papillose. |
oblong-conic, smooth, hairy, hairs papillose. |
Spores | 18–35 µm. |
14–17(–20) µm. |
Specialized | asexual reproduction by gemmae on leaves. |
asexual reproduction absent. |
Orthotrichum lyellii |
Orthotrichum shawii |
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Habitat | Trees, covering trunks to 10 m, boulders | Tree trunks |
Elevation | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) | elevation unknown |
Distribution |
AK; CA; ID; OR; WA; BC; Mexico (Baja California); Europe; n Africa
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CA; Europe |
Discussion | Orthotrichum lyellii is easily distinguished from all other species of the genus by its dioicous sexual condition, the male plants being slightly smaller and having numerous and conspicuous perigonia. In addition, the 16, slenderly acuminate, whitish exostome teeth and 16 endostome segments are characteristic. Sterile plants usually have abundant laminal gemmae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum shawii is similar to O. rupestre but distinguished by the 16 short, recurved exostome teeth and nearly smooth capsules, while O. rupestre has erect or spreading exostome teeth and variable capsule ribbing. Orthotrichum shawii was collected in California once without exact locality in the 1860s (Bolander 481780, NY). Stomata are found in the mid and distal capsule, the exostome teeth are irregularly divided, and the endostome segments are very short and hyaline. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 59. | FNA vol. 28, p. 66. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | Dorcadion lyellii, O. lyellii var. howei, O. lyellii var. papillosum, O. lyellii var. pringlei, O. lyellii subsp. strictum, O. lyellioides, O. menziesii, O. papillosum, O. papillosum subsp. strictum, O. pringlei | |
Name authority | Hooker & Taylor: Muscol. Brit., 76, plate 22 [upper left]. (1818) | Wilson: in W. P. Schimper, Musci Eur. Nov. 1 – 2(1): 1. (1864) |
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