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Lyell's bristle-moss, Lyell's orthotrichum moss

pylais' orthotrichum moss

Habit Plants to 13 cm. Plants to 3 cm, light green to olive green.
Stem

leaves erect-spreading to loosely erect, ± contorted when dry, lanceolate to linear-lanceolate, 2.5–6.5 mm;

margins plane, entire;

apex narrowly acute to acuminate;

basal laminal cells narrowly elongate, walls thick, ± nodose;

distal cells 7–14 µm, 1-stratose, papillae 1–3 per cell, conic, large.

leaves stiff, erect-appressed when dry, ovate-lanceolate to lanceolate, 2–3 mm;

margins revolute, entire;

apex broadly acute to narrowly obtuse, often bluntly acuminate;

basal laminal cells rectangular-elongate, walls thick, ± nodose;

distal cells 9–16 µm, 1-stratose, papillae 1–4 per cell, 2-fid, small.

Seta

0.8–2.7 mm.

2–3 mm.

Sexual condition

dioicous.

gonioautoicous.

Capsule

immersed, emergent, or rarely exserted, oblong-cylindric, 1.7–3 mm, strongly 8-ribbed to mid capsule or entire length;

stomata superficial;

peristome double;

prostome absent;

exostome teeth 16, reflexed-recurved, acuminate, densely papillose;

endostome segments 16, well developed, usually present when capsule is old and dry, thick, stout, of 2 rows of cells, narrower than exostome teeth, yellowish brown, coarsely papillose.

emergent or exserted, oblong to cylindric, 1.2–2.5 mm, slightly 8-ribbed distally;

exothecial cells differentiated to mid capsule;

stomata superficial;

peristome double or single;

prostome occasionally present, small;

exostome teeth 16, sometimes connate in 8 pairs, reflexed-recurved, densely papillose, not fenestrate, not cancellate;

endostome segments often absent, occasionally rudimentary, or 8, well developed, usually present in at least some capsules, narrow, of 1 row of cells, smooth or finely papillose.

Calyptra

conic-oblong, smooth, very hairy, hairs papillose.

oblong to oblong-conic, smooth, sparsely hairy, hairs smooth.

Spores

18–35 µm.

16–27 µm.

Specialized

asexual reproduction by gemmae on leaves.

asexual reproduction absent.

Orthotrichum lyellii

Orthotrichum pylaisii

Habitat Trees, covering trunks to 10 m, boulders Rock near oceans or alpine or arctic zones, on Salix or Alnus, bone, in or near bird rookeries, granitic boulders in coniferous forests
Elevation low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) low to high elevations (100- 3000 m) (low to high elevations (300- 9800 ft))
Distribution
from FNA
AK; CA; ID; OR; WA; BC; Mexico (Baja California); Europe; n Africa
[WildflowerSearch map]
from FNA
AK; CA; ID; MT; WA; WY; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; n Europe; n Asia
Discussion

Orthotrichum lyellii is easily distinguished from all other species of the genus by its dioicous sexual condition, the male plants being slightly smaller and having numerous and conspicuous perigonia. In addition, the 16, slenderly acuminate, whitish exostome teeth and 16 endostome segments are characteristic. Sterile plants usually have abundant laminal gemmae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Orthotrichum pylaisii is distinguished by the ribbed capsules with reflexed exostome teeth and a rudimentary endostome, along with superficial stomata and large, ovate-lanceolate leaves. The capsule is usually exserted in North American populations; however, in European populations it is sometimes only emergent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 59. FNA vol. 28, p. 64.
Parent taxa Orthotrichaceae > Orthotrichum Orthotrichaceae > Orthotrichum
Sibling taxa
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
Synonyms Dorcadion lyellii, O. lyellii var. howei, O. lyellii var. papillosum, O. lyellii var. pringlei, O. lyellii subsp. strictum, O. lyellioides, O. menziesii, O. papillosum, O. papillosum subsp. strictum, O. pringlei O. arcticum, O. arcticum subsp. blyttii, O. arcticum var. groenlandicum, O. blyttii, O. blyttii subsp. arcticum, O. blyttii var. arcticum, O. blyttii var. groenlandicum, O. blyttii subsp. microblephare, O. blyttii var. microblephare, O. groenlandicum, O. idaohense, O. pylaisii subsp. arcticum, O. pylaisii subsp. groenlandicum, O. pylaisii subsp. microblephare, O. rhabdophorum
Name authority Hooker & Taylor: Muscol. Brit., 76, plate 22 [upper left]. (1818) Bridel: Bryol. Univ. 1: 722. (1826)
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