Orthotrichum laevigatum |
Orthotrichum stellatum |
|
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orthotrichum moss |
stellate orthotrichum moss |
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Habit | Plants (0.5–)2(–4.3) cm, light green to olive green. | Plants 0.3–0.8 cm. |
Stem | leaves stiff, erect-appressed, slightly secund when dry, oblong-lanceolate, lanceolate, or ovate-lanceolate, 2–4.3 mm; margins revolute from base to near apex, entire; apex obtuse, bluntly or narrowly acute, or gradually acuminate; basal laminal cells rectangular to elongate, grading to elliptic-quadrate, walls thick, nodose; distal cells 8–12 µm, 1-stratose, papillae 1–3 per cell, conic or 2-fid, low. |
leaves stiff, loosely erect-appressed when dry, lanceolate to oblong-lanceolate, 1.8–2.2 mm; margins recurved to just below apex, entire; apex narrowly obtuse to bluntly acute, not incurved when dry; basal laminal cells rectangular, walls thin, nodose toward costa; distal cells 8–13 µm, 1-stratose, lumina irregular due to uneven wall thickenings, smooth or papillae 1–3 per cell, conic, small. |
Seta | 1.5–4 mm. |
0.4–1 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | fully exserted, cylindric, 1.4–2.6 mm, smooth or sometimes slightly plicate distally; exothecial cells homogeneous; stomata superficial; peristome double; prostome absent or rudimentary; exostome teeth 16, erect, papillose; endostome segments 8, occasionally rudimentary, usually absent in open capsules, of 1 row of cells, smooth. |
emergent, oblong-cylindric and strangulate when dry, 0.4–1 mm, 8-ribbed when dry, ribs thick, broad, strongly constricted below mouth; stomata immersed, mid capsule, 1/2 to completely covered by subsidiary cells, cells projecting horizontally over stomata, inner walls thickened; peristome double; prostome absent; exostome teeth 8, reflexed, finely and evenly papillose; endostome segments 8, well developed, of 1 or 2 rows of cells, smooth. |
Calyptra | conic-oblong, smooth, sparsely hairy, hairs finely papillose. |
oblong, smooth, naked or sparsely hairy, hairs smooth. |
Spores | 10–15 µm. |
13–18 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae on abaxial surface of leaves. |
Vaginula | with hairs absent. |
|
Orthotrichum laevigatum |
Orthotrichum stellatum |
|
Habitat | Non-calcareous boulders in mesic, open coniferous forests, base of deciduous trees | Hardwood trunks in moist forests, with Orthotrichum ohioense in mesic, shaded stream edges |
Elevation | moderate to high elevations (500-3000 m) (moderate to high elevations (1600-9800 ft)) | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Europe; e Asia (Japan)
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AR; DE; GA; IA; IL; IN; KY; MA; MD; ME; MI; MO; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; VA; VT; WV; NB; NF; NS; ON
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Discussion | Orthotrichum laevigatum is part of a complex group of intergrading taxa that generally are characterized by exserted, cylindric capsules with 16 erect or reflexed exostome teeth and a poorly developed endostome. Gametophytically, robust plants have stiff, erect leaves and large, blunt papillae. This complex is treated here as three species: O. holzingeri, a distinct, highly specialized species of seasonally wet rock with smooth exostome teeth, smooth laminal cells, smooth calyptra, and slightly ribbed capsule as distinguishing features; O. laevigatum, distinguished by smooth capsule and erect, papillose exostome teeth; and O. pylaisii, distinguished by slightly 8-ribbed capsule and reflexed-recurved exostome teeth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum stellatum is distinguished by the strongly ribbed, strangulate capsules with well-developed endostomes of eight segments. In comparison, O. ohioense has weakly ribbed capsules that are not constricted below the mouth and an endostome that is lacking or rudimentary. From O. pumilum, the narrowly lanceolate leaves and small laminal cells with unevenly incrassate walls differentiate O. stellatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 57. | FNA vol. 28, p. 68. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | O. hainesiae, O. kingianum, O. laevigatum var. kingianum, O. lonchothecium, O. macounii, O. macounii var. lonchothecium, O. raui, O. roellii, O. roellii subsp. schlotthaueri, O. roellii subsp. stenocarpum, O. schlotthaueri, O. speciosum var. brevicaule, O. speciosum var. hainesiae, O. speciosum subsp. laevigatum, O. speciosum var. polyanthum, O. speciosum var. polycarpum, O. speciosum var. raui | O. braunii, O. strangulatum var. stellatum |
Name authority | J. E. Zetterstedt: Öfvers. Kongl. Vetensk.-Akad. Förh. 19: 363. (1862) | Bridel: Bryol. Univ. 1: 274. (1826) |
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