Orthotrichum laevigatum |
Orthotrichum pumilum |
|
---|---|---|
orthotrichum moss |
low bristle moss, orthotrichum moss |
|
Habit | Plants (0.5–)2(–4.3) cm, light green to olive green. | Plants to 0.5 cm. |
Stem | leaves stiff, erect-appressed, slightly secund when dry, oblong-lanceolate, lanceolate, or ovate-lanceolate, 2–4.3 mm; margins revolute from base to near apex, entire; apex obtuse, bluntly or narrowly acute, or gradually acuminate; basal laminal cells rectangular to elongate, grading to elliptic-quadrate, walls thick, nodose; distal cells 8–12 µm, 1-stratose, papillae 1–3 per cell, conic or 2-fid, low. |
leaves stiff, loosely erect-appressed when dry, ovate-lanceolate to lanceolate, 1.8–2.8 mm; margins revolute nearly to apex, entire; apex sharply to slenderly acute, usually apiculate, rarely obtuse, not carinate, not incurved when dry; basal laminal cells rectangular, grading to quadrate at margin, walls thin, not nodose; distal cells 14–20 µm, 1-stratose, lumina smooth, rounded, smooth or papillae 1 or 2 per cell, conic, small. |
Seta | 1.5–4 mm. |
to 0.5 mm. |
Sexual condition | gonioautoicous. |
autoicous. |
Capsule | fully exserted, cylindric, 1.4–2.6 mm, smooth or sometimes slightly plicate distally; exothecial cells homogeneous; stomata superficial; peristome double; prostome absent or rudimentary; exostome teeth 16, erect, papillose; endostome segments 8, occasionally rudimentary, usually absent in open capsules, of 1 row of cells, smooth. |
immersed to 3/4 emergent, oblong to oblong-cylindric, 1.2–1.5 mm, strongly 8-ribbed, sometimes constricted below mouth when dry; stomata immersed, mid capsule, not or to 1/2 covered by subsidiary cells, cells projecting or not, inner walls thickened or not; peristome double; prostome absent; exostome teeth 8, rarely splitting to 16, reflexed, finely and evenly papillose; endostome segments 8, not well developed, of 1 or 2 rows of cells, smooth. |
Calyptra | conic-oblong, smooth, sparsely hairy, hairs finely papillose. |
oblong, smooth, sparsely hairy or rarely naked, hairs smooth. |
Spores | 10–15 µm. |
14–21 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction occasional, by gemmae on leaves. |
Vaginula | with hairs absent. |
|
Orthotrichum laevigatum |
Orthotrichum pumilum |
|
Habitat | Non-calcareous boulders in mesic, open coniferous forests, base of deciduous trees | Trunks, lower branches, base of deciduous trees, coniferous trees, crevices of rock, open, hardwood forests, trees along roadsides |
Elevation | moderate to high elevations (500-3000 m) (moderate to high elevations (1600-9800 ft)) | low to moderate elevations (10-1000 m) (low to moderate elevations (0-3300 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Europe; e Asia (Japan)
|
AZ; CA; CO; DC; IA; ID; IL; KS; MA; MD; MI; MN; MO; MS; MT; ND; NE; NH; NJ; NM; NY; OH; OK; PA; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; AB; MB; ON; SK; Europe; n Africa
|
Discussion | Orthotrichum laevigatum is part of a complex group of intergrading taxa that generally are characterized by exserted, cylindric capsules with 16 erect or reflexed exostome teeth and a poorly developed endostome. Gametophytically, robust plants have stiff, erect leaves and large, blunt papillae. This complex is treated here as three species: O. holzingeri, a distinct, highly specialized species of seasonally wet rock with smooth exostome teeth, smooth laminal cells, smooth calyptra, and slightly ribbed capsule as distinguishing features; O. laevigatum, distinguished by smooth capsule and erect, papillose exostome teeth; and O. pylaisii, distinguished by slightly 8-ribbed capsule and reflexed-recurved exostome teeth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum pumilum usually can be distinguished by the strongly ribbed capsules, reflexed peristome teeth, apiculate leaves with cells larger than 14 µm, and papillae that are very small or absent. In eastern North America, this taxon is uniform and non-variable, but in the West it is much more variable. Many western specimens have capsules that are oblong-cylindric, often strangulate, over 1/2 emergent, and gradually contracted to the seta. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 57. | FNA vol. 28, p. 63. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | O. hainesiae, O. kingianum, O. laevigatum var. kingianum, O. lonchothecium, O. macounii, O. macounii var. lonchothecium, O. raui, O. roellii, O. roellii subsp. schlotthaueri, O. roellii subsp. stenocarpum, O. schlotthaueri, O. speciosum var. brevicaule, O. speciosum var. hainesiae, O. speciosum subsp. laevigatum, O. speciosum var. polyanthum, O. speciosum var. polycarpum, O. speciosum var. raui | O. affine var. fallax, O. brachytrichum, O. fallax, O. fallax var. truncatulum, O. pumilum var. americanum, O. pumilum subsp. fallax, O. pumilum var. fallax, O. schimperi, O. schimperi var. truncatulum, O. tenellum var. pumilum |
Name authority | J. E. Zetterstedt: Öfvers. Kongl. Vetensk.-Akad. Förh. 19: 363. (1862) | Swartz: Monthly Rev. 34: 538. (1801) |
Web links |
|