Orthotrichum laevigatum |
Orthotrichum exiguum |
|
---|---|---|
orthotrichum moss |
orthotrichum moss |
|
Habit | Plants (0.5–)2(–4.3) cm, light green to olive green. | Plants 0.3–1 cm. |
Stem | leaves stiff, erect-appressed, slightly secund when dry, oblong-lanceolate, lanceolate, or ovate-lanceolate, 2–4.3 mm; margins revolute from base to near apex, entire; apex obtuse, bluntly or narrowly acute, or gradually acuminate; basal laminal cells rectangular to elongate, grading to elliptic-quadrate, walls thick, nodose; distal cells 8–12 µm, 1-stratose, papillae 1–3 per cell, conic or 2-fid, low. |
leaves erect-appressed when dry, oblong-lanceolate to oblong, 0.7–1.3 mm; margins reflexed to broadly recurved, entire; apex rounded-obtuse; basal laminal cells elongate, walls thick, ± nodose; distal cells 7–13 µm, 1-stratose, papillae 1 or 2 per cell, conic or C-shaped, small. |
Seta | 1.5–4 mm. |
to 5 mm. |
Sexual condition | gonioautoicous. |
gonioautoicous. |
Capsule | fully exserted, cylindric, 1.4–2.6 mm, smooth or sometimes slightly plicate distally; exothecial cells homogeneous; stomata superficial; peristome double; prostome absent or rudimentary; exostome teeth 16, erect, papillose; endostome segments 8, occasionally rudimentary, usually absent in open capsules, of 1 row of cells, smooth. |
immersed or slightly emergent, ovate-oblong, widest at mouth, less than 1 mm, distinctly 8-ribbed to base when dry; stomata superficial; peristome double; prostome absent; exostome teeth 16, spreading or reflexed when dry, papillose to cross striolate proximally, papillose to vertically or reticulately striolate distally; endostome segments 8, well developed, usually present when capsule is old and dry, thick, delicate, of 2 rows of cells, as wide as exostome teeth, hyaline, granulate-papillose. |
Calyptra | conic-oblong, smooth, sparsely hairy, hairs finely papillose. |
short-conic, finely papillose, naked or hairs few near tip, hairs finely papillose. |
Spores | 10–15 µm. |
17–20 µm. |
Specialized | asexual reproduction absent. |
asexual reproduction by gemmae on leaves. |
Orthotrichum laevigatum |
Orthotrichum exiguum |
|
Habitat | Non-calcareous boulders in mesic, open coniferous forests, base of deciduous trees | Base and trunks of trees |
Elevation | moderate to high elevations (500-3000 m) (moderate to high elevations (1600-9800 ft)) | low to moderate elevations (10-200 m) (low to moderate elevations (0-700 ft)) |
Distribution |
AK; AZ; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; MB; NT; SK; YT; Europe; e Asia (Japan)
|
GA; NC; SC; TN; VA; e Asia |
Discussion | Orthotrichum laevigatum is part of a complex group of intergrading taxa that generally are characterized by exserted, cylindric capsules with 16 erect or reflexed exostome teeth and a poorly developed endostome. Gametophytically, robust plants have stiff, erect leaves and large, blunt papillae. This complex is treated here as three species: O. holzingeri, a distinct, highly specialized species of seasonally wet rock with smooth exostome teeth, smooth laminal cells, smooth calyptra, and slightly ribbed capsule as distinguishing features; O. laevigatum, distinguished by smooth capsule and erect, papillose exostome teeth; and O. pylaisii, distinguished by slightly 8-ribbed capsule and reflexed-recurved exostome teeth. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Orthotrichum exiguum is distinguished by the distal laminal cells with bulging walls and C-shaped papillae; clavate stems; papillose, quadrate basal cells; differentiated perichaetial leaves; and capsules that are widest at the mouth, with 16 reflexed exostome teeth and delicate hyaline endostome. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 28, p. 57. | FNA vol. 28, p. 55. |
Parent taxa | Orthotrichaceae > Orthotrichum | Orthotrichaceae > Orthotrichum |
Sibling taxa | ||
Synonyms | O. hainesiae, O. kingianum, O. laevigatum var. kingianum, O. lonchothecium, O. macounii, O. macounii var. lonchothecium, O. raui, O. roellii, O. roellii subsp. schlotthaueri, O. roellii subsp. stenocarpum, O. schlotthaueri, O. speciosum var. brevicaule, O. speciosum var. hainesiae, O. speciosum subsp. laevigatum, O. speciosum var. polyanthum, O. speciosum var. polycarpum, O. speciosum var. raui | |
Name authority | J. E. Zetterstedt: Öfvers. Kongl. Vetensk.-Akad. Förh. 19: 363. (1862) | Sullivant: in A. Gray, Manual ed. 2, 633. (1856) |
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