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orthotrichum moss

Hall's orthotrichum moss

Habit Plants 0.5–2.3 cm. Plants to 2.5 cm.
Stem(s)

leaves loosely erect-appressed and ± loosely twisted when dry, elongate- to ovate-lanceolate, 2–3.5 mm;

margins revolute to below apex, entire;

apex sharply acute, sometimes acuminate, cuspidate, or apiculate;

basal laminal cells long-rectangular to rectangular, walls thin, sometimes nodose at insertion;

distal cells 9–14 µm, 1-stratose, papillae 2 or 3 per cell, conic or 2-fid, large.

leaves loosely erect-appressed when dry, elongate-lanceolate to lanceolate, 1.7–3.5 mm;

margins revolute proximally, recurved to almost plane near apex, entire;

apex ± narrowly obtuse, rarely bluntly acute;

basal laminal cells rectangular to short-rectangular, grading to quadrate at margins, walls thin, not nodose;

distal cells 7–13 µm, 2-stratose, rarely with few 1-stratose areas, papillae 1–3 per cell, conic, small;

marginal cells mostly to completely 2-stratose.

Seta

to 1.4 mm.

0.5–1 mm.

Sexual condition

gonioautoicous.

gonioautoicous.

Capsule

1/2–3/4 emergent, sometimes almost exserted, oblong-ovate when mature, oblong to short-cylindric when old and dry, to 1.4 mm, strongly 8-ribbed 2/3 to entire length, constricted below mouth when dry;

stomata immersed, 1/2 covered by subsidiary cells, cells projecting, inner walls thickened;

peristome double;

prostome absent;

exostome teeth 8, reflexed, densely and evenly papillose, striate-reticulate distally;

endostome segments 8 or 16, well developed, of 1 or 2 rows of cells, smooth or finely roughened.

1/2 emergent when dry, immersed when moist, oblong or oblong-ovate, 1–1.8 mm, strongly 8-ribbed to mid capsule or entire length, rarely with 8 very short intermediate ribs, sometimes constricted below mouth when dry;

stomata immersed;

peristome double;

prostome present, occasionally rudimentary;

exostome teeth 8, sometimes irregularly split to 16, incurved becoming spreading or rarely reflexed when old, coarsely striate-reticulate or striate-papillose;

endostome segments 8, not well developed, of 1 row of cells, short, finely longitudinally striate.

Calyptra

oblong, smooth, ± hairy, hairs strongly papillose.

oblong, smooth, sparsely hairy, hairs papillose.

Spores

10–15 µm.

10–17 µm.

Specialized

asexual reproduction on leaves.

asexual reproduction absent.

Orthotrichum alpestre

Orthotrichum hallii

Habitat Rock, trees, base of trees, moist crevices of large boulders, moist pine and deciduous forests Rock, usually limestone or calcareous sandstone, granite, quartzite, basalt, trunks of deciduous trees, open pine forests, spruce-fir forests, deciduous scrub oak-maple forests, vertical canyon walls, shaded cliff faces
Elevation high elevations (2100-3000 m) (high elevations (6900-9800 ft)) moderate to high elevations (200-3000 m) (moderate to high elevations (700-9800 ft))
Distribution
from FNA
AZ; CA; CO; ID; MI; MT; NM; NV; OR; SD; UT; WA; WY; AB; BC; NL; ON; YT; Asia; Greenland; Europe; Pacific Islands (New Zealand)
[WildflowerSearch map]
from FNA
AZ; CA; CO; ID; MT; NM; NV; SD; UT; WY; BC
Discussion

Orthotrichum alpestre is distinguished by the highly papillose distal laminal cells; slightly twisted, sharply pointed leaves; calyptra with papillose hairs; absence of prostome; and papillose or papillose-reticulate exostome. The endostome is always present in mature capsules and, because of enlarged cross walls, has a shiny appearance. The capsule is usually oblong and strangulate, although the shape depends on age. In some specimens, the capsule is short-cylindric and not much constricted below the mouth. The stomata are found just below mid capsule.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Orthotrichum hallii is distinguished by leaves that are completely or nearly 2-stratose in their distal portion, and emergent, oblong, 8-ribbed capsules. The peristome teeth vary from ridged-striate to papillose-striate; the proximal portions of the teeth are often coarsely papillose with the distal portions having striae mixed with papillae, and the papillae are always small and conic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 50. FNA vol. 28, p. 56.
Parent taxa Orthotrichaceae > Orthotrichum Orthotrichaceae > Orthotrichum
Sibling taxa
O. affine, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
Synonyms O. alpestre var. majus, O. alpestre var. watsonii, O. occidentale, O. watsonii
Name authority Hornschuch ex Bruch & Schimper: Bryol. Europ. 3: 75, plate 213. (1849) Sullivant & Lesquereux: in W. S. Sullivant, Icon. Musc., suppl., 63, plate 45. (1874)
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