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orthotrichum moss, wood bristle-moss

lanceolateleaf rock moss, showy bristle-moss

Habit Plants to 3 cm. Plants 0.8–5 cm.
Stem

leaves erect-appressed and straight when dry, ligulate, elongate- to ovate-lanceolate, 2–4 mm;

margins recurved-revolute to just below apex, entire;

apex acute;

basal laminal cells long-rectangular to elongate, walls thick, nodose;

distal cells 9–13 µm, 1-stratose, papillae 2 or 3 per cell, 2-fid, low.

leaves loosely erect, usually slightly contorted or curved when dry, lanceolate to narrowly lanceolate, 3–4 mm;

margins revolute nearly to apex, entire;

apex acuminate to narrowly acute;

basal laminal cells elongate, walls thick, nodose or wavy;

distal cells 7–13 µm, 1-stratose, papillae 1–3 per cell, conic or 2-fid, large.

Seta

1–2.2 mm.

1–3 mm.

Sexual condition

gonioautoicous.

gonioautoicous.

Capsule

1/2 emergent to short-exserted, long-cylindric when mature, cylindric and constricted along entire length when old and dry, 1.6–2.5 mm, strongly 8-ribbed 2/3 capsule length;

stomata superficial;

peristome double;

prostome absent;

exostome teeth 8, erect when mature, reflexed to reflexed-recurved when old, coarsely and obscurely papillose or sometimes papillose-reticulate, deeply perforate, cancellate;

endostome segments 8, rarely 16, well developed, usually present when capsule is old and dry, narrow, of 2 rows of cells, papillose.

exserted to rarely emergent, cylindric to ovate-cylindric when mature, 1.5–2.4 mm, slightly 8-ribbed to mid capsule;

stomata superficial;

peristome double;

prostome absent;

exostome teeth 8, separate to base, recurved, acute, coarsely papillose;

endostome segments 8, well developed, usually present when capsule is old and dry, thick, stout, of 2 rows of cells, narrower than exostome teeth, yellowish brown, coarsely papillose.

Calyptra

conic-oblong, smooth, hairs few, papillose.

conic-oblong, smooth, hairy, hairs finely papillose.

Spores

15–18 µm.

13–26 µm.

Specialized

asexual reproduction absent.

asexual reproduction absent.

Orthotrichum affine

Orthotrichum speciosum

Habitat Deciduous trees, shaded rock, shaded forests near stream beds, canyons, mountains Coniferous and deciduous trees, rock
Elevation low to high elevations (0-3000 m) (low to high elevations (0-9800 ft)) low to high elevations (10-2000 m) (low to high elevations (0-6600 ft))
Distribution
from FNA
CA; CO; ID; MT; OR; UT; WA; AB; BC; Europe; Asia; n Africa
[WildflowerSearch map]
from FNA
AK; CA; CO; ID; MI; MN; MT; NH; NV; NY; OR; UT; WA; WY; AB; BC; MB; NB; NL; NS; NT; NU; PE; QC; SK; YT; n Asia; Greenland; Europe; n Africa
[WildflowerSearch map]
Discussion

Orthotrichum affine, a species confined to western North America in the flora area, is best recognized by: superficial stomata; cylindric, strongly ribbed, almost exserted capsules; a well-developed but not stout endostome; and a calyptra that is only sparsely hairy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Orthotrichum speciosum is primarily western and Arctic in distribution; the plants are distinguished by their ribbed capsules, large size, and more slender leaf apices.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 50. FNA vol. 28, p. 67.
Parent taxa Orthotrichaceae > Orthotrichum Orthotrichaceae > Orthotrichum
Sibling taxa
O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. speciosum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
O. affine, O. alpestre, O. anodon, O. anomalum, O. bartramii, O. bolanderi, O. consimile, O. cupulatum, O. diaphanum, O. elegans, O. euryphyllum, O. exiguum, O. fenestratum, O. flowersii, O. gymnostomum, O. hallii, O. holzingeri, O. keeverae, O. kellmanii, O. laevigatum, O. lescurii, O. lyellii, O. norrisii, O. obtusifolium, O. ohioense, O. pallens, O. parvulum, O. pellucidum, O. praemorsum, O. pulchellum, O. pumilum, O. pusillum, O. pycnophyllum, O. pylaisii, O. rivulare, O. rupestre, O. shawii, O. shevockii, O. sordidum, O. spjutii, O. stellatum, O. stramineum, O. strangulatum, O. striatum, O. tenellum, O. underwoodii
Synonyms O. killiasii, O. speciosum subsp. killiasii, O. speciosum var. killiasii
Name authority Schrader ex Bridel: Muscol. Recent. 2(2): 22. (1801) Nees: in J. Sturm et al., Deutschl. Fl. 2 (17): 5. (1819)
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