Orthotrichum affine |
Orthotrichaceae |
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orthotrichum moss, wood bristle-moss |
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Habit | Plants to 3 cm. | Plants small to large, in tufts, cushions, or mats, dark green, reddish brown, or olive brown, dull. | ||||||||||||||||||||||||||||||||
Stem(s) | leaves erect-appressed and straight when dry, ligulate, elongate- to ovate-lanceolate, 2–4 mm; margins recurved-revolute to just below apex, entire; apex acute; basal laminal cells long-rectangular to elongate, walls thick, nodose; distal cells 9–13 µm, 1-stratose, papillae 2 or 3 per cell, 2-fid, low. |
erect-ascending and often 2-fid, or creeping, branches many, erect or ascending, simple or 2-fid; outer cell walls thick, central strand absent, inner cells uniform, walls thin. |
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Leaves | erect-appressed, crowded, spirally twisted, flexuose, crisped, or contorted when dry, erect-spreading to squarrose-recurved when moist, ovate-lanceolate, ligulate-oblong, or lanceolate-linear, ± channeled; base not decurrent (decurrent in Zygodon); margins usually plane to revolute, rarely involute to erect, entire or sometimes denticulate near apex; apex rounded-obtuse, acute, occasionally acuminate, apiculate, or awned; costa strong, ending near apex; alar cells rarely differentiated; basal laminal cells rectangular, elongate-linear, quadrate, rounded, or elliptic; distal cells rounded-hexagonal, rarely rectangular, usually small, papillae 1–4(–6) per cell, conic or 2-fid, rarely smooth or mammillose, walls often incrassate. |
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Seta | 1–2.2 mm. |
erect, dextrorse or sinistrorse, smooth or rarely rough. |
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Sexual condition | gonioautoicous. |
usually gonioautoicous or dioicous, rarely pseudautoicous or cladautoicous; perigonia terminal or lateral, sometimes occurring on dwarf male plants, budlike, large; perichaetia terminal, further branching occurring by innovations, perichaetial leaves sometimes larger than stem leaves. |
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Capsule | 1/2 emergent to short-exserted, long-cylindric when mature, cylindric and constricted along entire length when old and dry, 1.6–2.5 mm, strongly 8-ribbed 2/3 capsule length; stomata superficial; peristome double; prostome absent; exostome teeth 8, erect when mature, reflexed to reflexed-recurved when old, coarsely and obscurely papillose or sometimes papillose-reticulate, deeply perforate, cancellate; endostome segments 8, rarely 16, well developed, usually present when capsule is old and dry, narrow, of 2 rows of cells, papillose. |
erect, immersed, emergent, or exserted, ovate to cylindric-fusiform, symmetric, smooth or 8-ribbed, rarely 16-ribbed, sometimes constricted below mouth; exothecial cells rectangular to elliptic, often differentiated into bands; stomata superficial or immersed, usually below mid capsule, well developed; annulus poorly developed or absent; operculum convex to conic, rostrate; peristome double, single, rudimentary or rarely absent; prostome sometimes present, usually fragmentary; exostome teeth 16, usually connate in 8 pairs, erect, recurved, or reflexed, lanceolate, thick, densely papillose or striate; endostome segments 8 or 16 when present, hyaline, thin, linear-lanceolate, alternating with exostome teeth or connate to form rudimentary membrane, cilia absent. |
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Calyptra | conic-oblong, smooth, hairs few, papillose. |
mitrate or rarely cucullate, usually large and conspicuous, hairy or naked, plicate or not. |
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Spores | 15–18 µm. |
isosporous or anisosporous. |
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Specialized | asexual reproduction absent. |
asexual reproduction occasional, by gemmae. |
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Orthotrichum affine |
Orthotrichaceae |
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Habitat | Deciduous trees, shaded rock, shaded forests near stream beds, canyons, mountains | |||||||||||||||||||||||||||||||||
Elevation | low to high elevations (0-3000 m) (low to high elevations (0-9800 ft)) | |||||||||||||||||||||||||||||||||
Distribution |
CA; CO; ID; MT; OR; UT; WA; AB; BC; Europe; Asia; n Africa
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Nearly worldwide |
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Discussion | Orthotrichum affine, a species confined to western North America in the flora area, is best recognized by: superficial stomata; cylindric, strongly ribbed, almost exserted capsules; a well-developed but not stout endostome; and a calyptra that is only sparsely hairy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 22, species ca. 380 (9 genera, 71 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 28, p. 50. | FNA vol. 28, p. 37. | ||||||||||||||||||||||||||||||||
Parent taxa | Orthotrichaceae > Orthotrichum | |||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||
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Name authority | Schrader ex Bridel: Muscol. Recent. 2(2): 22. (1801) | Schimper | ||||||||||||||||||||||||||||||||
Web links |