Opuntia littoralis |
Opuntia ×columbiana |
|
---|---|---|
coastal prickly pear, prickly pear |
Columbia prickly-pear, grizzleybear pricklypear, grizzlybear prickly pear |
|
Habit | Shrubs, spreading to sprawling, forming large clumps, to 1 × 1–9 m. | Shrubs, low, forming mats, 10–30 cm. |
Stem | segments not disarticulating, green, flattened, elliptic to obovate to rhombic, 15–25(–40) × 6.5–14 cm, ± tuberculate, glabrous, usually glaucous; areoles 5–7(–8) per diagonal row across midstem segment, prominent, subcircular(-oval), 4–6 × 4–5 mm, enlarging in age; wool gray. |
segments easily detached to firmly attached, green, flattened, narrowly to broadly obovate, 3.5–9 × 2.5–5 cm, low tuberculate, glabrous; areoles 4–6(–7) per diagonal row across midstem segment, oval, 3.5 × 2.5 mm; wool white. |
Spines | 4–11 per areole, in most areoles, yellow with chalky white coat, to yellow with red-brown basal portions, aging reddish gray; erect ones terete, stout, straight; abaxial ones reflexed, shorter, to 12 mm; adaxial spines spreading, longest spines 20–40 mm. |
in most areoles, straight, acicular, terete; major spines 0–1 per areole, reflexed to porrect (rarely ascending), gray-white to brown, (15–)25–40(–60) mm; intermediate spines 0–3, reflexed, 12–20 mm; minor spines 2–4, reflexed, yellow-gray, 3–6 mm. |
Glochids | moderately dense in crescent at adaxial edge of areole, merging with subapical tuft when present, yellow to red-brown, to 5 mm. |
loosely packed, in small tuft or crescent at adaxial margin of areole, yellowish, inconspicuous, to 4 mm. |
Flowers | inner tepals yellow to dull red throughout, 35–45 mm; filaments yellow to orange-yellow; anthers yellow; style pink to red; stigma lobes yellow-green to green. |
inner tepals yellow throughout, 30–45 mm; filaments white to red; anthers yellow; style white; stigma lobes green. |
Fruits | dark red-purple throughout, obovoid, 35–50 × 30–35 mm, juicy, glabrous, spineless; areoles 22–36. |
rarely set, top-shaped to barrel-shaped, 15–20 × 15 mm, glabrous; areoles 10–15, most bearing 3–12 spines or distal spines bearing 1–3 spines. |
Seeds | gray, subcircular, warped, 3–4.5 mm diam.; girdle protruding 0.5 mm. |
tan, large, 5–7 mm; girdle protruding 1–1.5 mm. |
2n | = 66. |
= 66. |
Opuntia littoralis |
Opuntia ×columbiana |
|
Phenology | Flowering spring (Apr–May). | Flowering summer (late Jun-early Jul). |
Habitat | Coastal sage scrub, chaparral | Dark basaltic cliffs and derived sands |
Elevation | 10-400 m (0-1300 ft) | 100-800 m (300-2600 ft) |
Distribution |
Calif (including Channel Islands); Mexico (Baja California)
|
ID; OR; WA; BC
|
Source | FNA vol. 4. | FNA vol. 4, p. 146. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Opuntia | Cactaceae > subfam. Opuntioideae > Opuntia |
Sibling taxa | ||
Synonyms | O. engelmannii var. littoralis, O. lindheimeri var. littoralis, O. occidentalis var. littoralis, O. semispinosa | O. erinacea var. columbiana |
Name authority | (Engelmann) Cockerell: Bull. S. Calif. Acad. Sci. 4: 15. (1905) | Griffiths: Bull. Torrey Bot. Club 43: 523. (1916) |
Web links |