Opuntia littoralis |
Opuntia macrocentra |
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coastal prickly pear, prickly pear |
black-spine pricklypear, nopal violaceo, purple pricklypear |
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Habit | Shrubs, spreading to sprawling, forming large clumps, to 1 × 1–9 m. | Shrubs, erect to decumbent, to 1 m. |
Stem | segments not disarticulating, green, flattened, elliptic to obovate to rhombic, 15–25(–40) × 6.5–14 cm, ± tuberculate, glabrous, usually glaucous; areoles 5–7(–8) per diagonal row across midstem segment, prominent, subcircular(-oval), 4–6 × 4–5 mm, enlarging in age; wool gray. |
segments not easily detached, purple (particularly under stress) to green with purple near areoles and margins of stem segment, flattened, broadly obovate to subcircular, thickish, 7–20 × 6–18 cm; areoles 6–8(–10) per diagonal row across midstem segment, elliptic to circular, 3–7 × 2.5–5 mm; wool tan to whitish, aging black. |
Spines | 4–11 per areole, in most areoles, yellow with chalky white coat, to yellow with red-brown basal portions, aging reddish gray; erect ones terete, stout, straight; abaxial ones reflexed, shorter, to 12 mm; adaxial spines spreading, longest spines 20–40 mm. |
0 or 1–15+ per areole, few and at distal areoles or many and on most areoles, usually erect to spreading, appearing unruly, sometimes deflexed, straight or curving, acicular, often flexible, subterete to flattened basally; longer ones reddish brown to ± black (rarely yellow to red in w Texas), or partly to wholly white, 30–120(–170) mm; reflexed spine rarely present, 1 in some areoles, short, whitish. |
Glochids | moderately dense in crescent at adaxial edge of areole, merging with subapical tuft when present, yellow to red-brown, to 5 mm. |
dense in crescent at adaxial edge of areole and well-developed subapical tuft, reddish yellow, aging brown, 2–3(–6) mm. |
Flowers | inner tepals yellow to dull red throughout, 35–45 mm; filaments yellow to orange-yellow; anthers yellow; style pink to red; stigma lobes yellow-green to green. |
inner tepals yellow with red basal portions, obovate-apiculate, 25–40 mm; filaments and anthers yellowish; style cream; stigma lobes green. |
Fruits | dark red-purple throughout, obovoid, 35–50 × 30–35 mm, juicy, glabrous, spineless; areoles 22–36. |
red to purplish, obovoid to barrel-shaped, 25–40 × 20–23 mm, fleshy or ± juicy, glabrous, spineless; umbilicus 8–10 mm deep; areoles 22–44. |
Seeds | gray, subcircular, warped, 3–4.5 mm diam.; girdle protruding 0.5 mm. |
yellowish, suborbicular to reniform, angled, 5–7 × 3.5–5 mm, sides flattened; girdle protruding 0.8–1.2 mm. |
2n | = 66. |
= 22, 44. |
Opuntia littoralis |
Opuntia macrocentra |
|
Phenology | Flowering spring (Apr–May). | Flowering spring (Mar–Jun). |
Habitat | Coastal sage scrub, chaparral | Desert uplands, grasslands, oak woodlands, sandy desert flats, rocky hills and valleys |
Elevation | 10-400 m (0-1300 ft) | 900-1600 m (3000-5200 ft) |
Distribution |
Calif (including Channel Islands); Mexico (Baja California)
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AZ; NM; TX; Mexico (Chihuahua, Coahuila, Sonora)
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Discussion | Opuntia macrocentra var. minor is represented by relatively short and compact shrubs with a glochid pattern of a dense crescent in the adaxial edge of the areoles like the species and a much taller tuft of spreading glochids. This variety, which grows along the western side of the Rio Grande in Big Bend, Texas, appears best interpreted as a tetraploid hybrid between tetraploid putative parents, Opuntia macrocentra and O. tortispina. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4. | FNA vol. 4. |
Parent taxa | Cactaceae > subfam. Opuntioideae > Opuntia | Cactaceae > subfam. Opuntioideae > Opuntia |
Sibling taxa | ||
Synonyms | O. engelmannii var. littoralis, O. lindheimeri var. littoralis, O. occidentalis var. littoralis, O. semispinosa | O. violacea, O. violacea var. castetteri, O. violacea var. macrocentra |
Name authority | (Engelmann) Cockerell: Bull. S. Calif. Acad. Sci. 4: 15. (1905) | Engelmann: Proc. Amer. Acad. Arts 3: 292. (1856) |
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