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Habit Herbs, perennial, or shrubs, [epiphytes, lianas, or trees].
Leaves

opposite or whorled, [alternate];

stipules present.

Flowers

primarily protogynous, actinomorphic and 4-merous, or zygomorphic and 2-merous;

stamens 2 times as many, or as many, as sepals;

pollen shed in monads.

Fruits

indehiscent, either a fleshy berry or a dry capsule, covered with stiff, hooked hairs.

Seeds

1–500, without hairs or wings.

Onagraceae tribe Circaeeae

Distribution
North America; Mexico; Central America; South America; West Indies (Hispaniola); Eurasia; n Africa; Pacific Islands (New Zealand, Society Islands)
Discussion

Genera 2, species 117 (2 genera, 4 species, including 1 hybrid, in the flora).

All previous classification systems have placed Circaea and Fuchsia into different tribes, based on their morphological and geographical differences. Molecular analyses place these genera into a single clade (C. J. Bult and E. A. Zimmer 1993; E. Conti et al. 1993; R. A. Levin et al. 2003, 2004; V. S. Ford and L. D. Gottlieb 2007) that is as or more strongly supported than are other clades. The two genera share the feature of indehiscent fruits, expressed in Fuchsia as fleshy berries and in Circaea as dry fruits covered with hooklike hairs; nonhomologous indehiscent fruits also occur in Onagreae. The only occurrences of protogyny in the family occur in these two genera (not in all species of either, P. H. Raven 1979).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae
Subordinate taxa
Synonyms Fuchsieae de
Name authority Dumortier: FFl. Belg., 88. (1827)
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