Oenothera xylocarpa |
Oenothera pubescens |
|
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wood fruit evening primrose, woodyfruit evening primrose |
pubescent evening primrose, silky evening primrose, South American evening-primrose |
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Habit | Herbs perennial, acaulescent, densely short-hirsute, also sometimes sparsely long-hirsute distally; from a thick, fleshy taproot. | Herbs annual or biennial, densely to sparsely strigillose, sometimes also villous and glandular puberulent distally; from a taproot. |
Stems | unbranched or with branched central stem and ascending to decumbent lateral branches arising from rosette, 5–50(–80) cm. |
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Leaves | in a basal rosette, 2.6–4.2(–6.2) × 1.4–4.2 cm; petiole 2.5–9(–11.5) cm; blade usually oblanceolate to obovate, sometimes suborbiculate, margins dentate, pinnately lobed, lateral lobes oblong to lanceolate, often absent or reduced to only a few lobes toward terminal lobe, base rounded to cordate. |
in a basal rosette and cauline, basal 5–14 × 0.5–2.5 cm, cauline 2–8 × 0.5–2.5 cm; blade narrowly oblanceolate to narrowly elliptic or narrowly oblong, margins usually dentate to deeply lobed; bracts spreading, flat. |
Flowers | usually 1–3, rarely more, opening per day near sunset; buds erect, quadrangular, without free tips; floral tube 27–45(–55) mm; sepals 25–30 mm; petals intensely yellow, fading deep salmon red, obcordate, 25–38 mm; filaments 17–23 mm, anthers 7–10 mm; style 44–65(–80) mm, stigma somewhat exserted beyond anthers at anthesis. |
usually 1 opening per day near sunset; buds with free tips erect, 0.1–1 mm; floral tube erect, becoming recurved and nodding, then erect again just before anthesis, 15–50 mm; sepals 5–25 mm; petals yellow, fading reddish orange, broadly obovate to obcordate, 5–25(–35) mm; filaments 6–18 mm, anthers (2–)3–9 mm, pollen ca. 50% fertile; style 20–60 mm, stigma surrounded by or slightly exserted beyond anthers at anthesis. |
Capsules | moderately thin and flexible, lanceoloid, falcate or sigmoid, often contorted and twisted, 4-angled, 35–90 × 7–11 mm, gradually tapering to a long, slender, sterile apex, 10–30(–40) mm, valves conspicuously wrinkled, dehiscent 2/3–3/4 their length; sessile. |
cylindrical, sometimes slightly enlarged distally, 20–45 ×2–4 mm. |
Seeds | numerous, in 1 row per locule, often forming 2 rows near base of capsule, obovoid, 2.4–3.2 × 1.3–1.7 mm, surface coarsely rugose. |
brown, sometimes dark-flecked, 0.9–1.5 × 0.6–1 mm. |
2n | = 14. |
= 14. |
Oenothera xylocarpa |
Oenothera pubescens |
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Phenology | Flowering Jun–Jul(–Aug). | Flowering (Feb–)Apr–Sep(–Oct). |
Habitat | Open meadows, flats or slopes on loose granitic gravel, sand, or pumice in Pinus jeffreyi forests with Artemisia tridentata, or in Pinus contorta subsp. murrayana and Abies magnifica forests. | Open sites in montane habitats. |
Elevation | 2200–3100 m. (7200–10200 ft.) | (1300–)1500–2500(–3100) m. ((4300–)4900–8200(–10200) ft.) |
Distribution |
CA; NV |
AZ; NM; TX; Mexico; West Indies; Central America (Guatemala); South America (Colombia, Ecuador, Peru)
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Discussion | Oenothera xylocarpa is known from three disjunct areas in California and adjacent Nevada: Mount Rose, Washoe County, Nevada; southern Sierra Nevada, southwestern Mono County, California, from the vicinity of Crestview south to Casa Diablo; and the area in the southern Sierra Nevada bounded by Horseshoe and Big Whitney meadows to the east and north, and Casa Vieja and Volcano Meadows to the south and west, west-central Inyo and eastern Tulare counties, California. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera pubescens is a PTH species and forms a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous (W. Dietrich and W. L. Wagner 1988). Oenothera pubescens has been collected once in California in 1884 (Newberry Springs, San Bernardino County), where it was temporarily introduced or a natural occurrence that was extirpated. Collections from west Texas (Brewster, Jeff Davis, and Presidio counties) have been made since 1990 and a few others collected earlier were misidentified as O. laciniata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Anogra xylocarpa | Anogra amplexicaulis, O. amplexicaulis, O. laciniata subsp. pubescens, Var. o. var. o., O. nyctaginiifolia, O. stuebelii, Raimannia colimae, R. confusa |
Name authority | Coville: Contr. U.S. Natl. Herb. 4: 105, plate 8. (1893) | Willdenow ex Sprengel: Syst. Veg. 2: 229. (1825) |
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