Oenothera villosa subsp. villosa
Onagraceae tribe Onagreae
hairy evening-primrose
grayish green to dull green, blade margins conspicuously dentate, sometimes sinuate-dentate proximally, venation prominent, especially abaxially, usually pale green, rarely red.
alternate or basal;
stipules absent.
relatively dense, apex truncate, internodes in fruit conspicuously shorter than capsules.
sepals green to yellowish green.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Oenothera villosa subsp. villosa
Onagraceae tribe Onagreae
Subspecies villosa appears to be largely a taxon of the Great Plains that has subsequently, both naturally and with human assistance, spread to the north and east, primarily in historical times. It grows in a variety of habitats, primarily prairies, along streams or lakes, open woodlands, old fields, and other disturbed sites, and is widely naturalized in Asia, Europe, southern South America, and South Africa.
Oenothera strigosa (Rydberg) Mackenzie & Bush var. depressa (Greene) R. R. Gates is an illegitimate name, as is O. salicifolia Desfontaines ex G. Don 1837, not Lehmann 1824 and not Desfontaines ex Seringe 1825, and these names pertain here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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