Oenothera tubicula subsp. tubicula
Onagraceae tribe Onagreae
1–many, unbranched to densely branched, decumbent to erect, 4–53 cm.
0.7–4.6 × 0.1–1.2 cm, sometimes fascicles of small leaves 0.2–1.5 cm present in non-flowering axils;
petiole 0–0.2 cm;
blade linear to ovate or obovate, base attenuate, margins entire, apex acute.
alternate or basal;
stipules absent.
usually several per stem opening per day near sunrise;
buds with free tips 0.5–2 mm;
floral tube 5–25(–33) mm, funnelform in distal 1/2 or more;
sepals 3–13 mm;
petals yellow, fading pale pink or pale purple, 5–20(–25) mm;
filaments 1–6 mm, anthers 2–7 mm, pollen 85–100% fertile;
style 9–30(–40) mm, stigma yellow, quadrangular, usually exserted beyond anthers.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
8–20 × 1.5–2.5 mm, hard, promptly dehiscent throughout their length.
obovoid, 1–1.4 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Oenothera tubicula subsp. tubicula
Onagraceae tribe Onagreae
Subspecies tubicula is known from Guadalupe County, New Mexico, south in the western side of the Pecos River drainage to western Texas, where it occurs from Culberson County east to Howard County, thence south through Brewster, Presidio, and Terrell counties, and probably most of central Coahuila, to northern Zacatecas, southwestern Nuevo León, and southwestern Tamaulipas.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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