Calylophus tubicula(synonym of Oenothera tubicula) |
Oenothera sinuosa |
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wavy-leaf gaura, wavyleaf beeblossom |
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Habit | Herbs perennial, usually glabrous, sometimes strigillose and villous, hairs erect; from a woody taproot but spreading by rhizomes (forming extensive colonies). | |
Stems | erect, branched below and just above ground, branched also proximal to inflorescences, 40–120(–250) cm. |
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Leaves | in a basal rosette and cauline, (1–)3–11 × (0.1–)0.5–2 cm, blade linear to narrowly oblanceolate, margins usually sparsely sinuate-dentate, rarely subentire, often undulate. |
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Inflorescences | stout. |
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Flowers | 4-merous, zygomorphic, opening near sunset; floral tube 2.5–5 mm; sepals 7–14 mm; petals white, fading pink to red, slightly unequal, elliptic, 7–15 mm; stamens presented in lower 1/2 of flower, filaments 5–11 mm, lanate at very base, anthers 3–5 mm, pollen 90–100% fertile; style 12–19 mm, stigma exserted beyond anthers at anthesis. |
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Capsules | narrowly ovoid, narrowly 4-winged or 4-angled, 8–15 × 1.5–3.5 mm, abruptly constricted to a long, sterile stipe 2–8 mm. |
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Seeds | (1 or)2–4, light to reddish brown, 2–3 × 1–1.5 mm. |
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2n | = 28. |
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Calylophus tubicula |
Oenothera sinuosa |
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Phenology | Flowering Apr–Aug. | |
Habitat | Flats and washes in light sandy loam. | |
Elevation | 0–300(–1300) m. (0–1000(–4300) ft.) | |
Distribution |
sw United States; sc United States; n Mexico |
AL; AR; CA; FL; GA; MO; NY; OK; TX [Introduced in Europe (Italy), s Africa]
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Discussion | Subspecies 2 (1 in the flora). H. F. Towner (1977) found that Oenothera tubicula is self-incompatible and diurnal with opening times just prior to sunrise. It occurs primarily on limestone soil in arid lowlands, but occasionally in montane areas, from Guadalupe County, New Mexico, south to western Texas, northeast to Howard County, Texas, and south to northern Zacatecas, south-central Nuevo León, and southwestern Tamaulipas, 600–1800 m. Subspecies strigulosa (Towner) W. L. Wagner & Hoch is known only from rocky, open sites and canyons in relatively montane areas, sometimes in pine forests in southernmost Coahuila, south-central Nuevo León, and southeastern Tamaulipas, from 1500 to 2300 m. It differs in being strigillose on the ovary and distally on stems, leaves linear to narrowly lanceolate, and the petals fading red or purple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
P. H. Raven and D. P. Gregory (1972[1973]) determined Oenothera sinuosa to be self-incompatible. Oenothera sinuosa is endemic to Oklahoma and Texas and is escaped or naturalized in Alabama, Arkansas, California (where found to 1300 m), Florida, Georgia, Missouri, and New York. Oenothera sinuosa is potentially a noxious weed due to the aggressive rhizomatous habit, but is somewhat limited by its self-incompatibility. Molecular data (G. D. Hoggard et al. 2004) are consistent with the hypothesis that the allotetraploid (2n = 28) O. sinuosa arose by interspecific hybridization of two species within subsect. Stipogaura as suggested by P. H. Raven and D. P. Gregory (1972[1973]). The molecular data indicate that the pistillate parent came from O. calcicola or a close relative, while the staminate parent originated from the lineage that gave rise to O. cinerea and O. filipes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Calylophus tubiculus, Galpinsia tubicula, O. hartwegii var. tubicula | Gaura sinuata |
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 71. (1852) | W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 214. (2007) |
Web links |