Calylophus tubicula(synonym of Oenothera tubicula) |
Oenothera humifusa |
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seabeach evening-primrose |
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Habit | Herbs annual or short-lived perennial, densely strigillose, sometimes also villous, also becoming glandular puberulent distally. | |
Stems | erect to decumbent, much branched, 10–50(–90) cm. |
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Leaves | in a basal rosette and cauline, basal 4–8 × 0.7–1 cm, cauline 1–7 × 0.3–1.5 cm; blade usually grayish green, narrowly oblong to narrowly elliptic or narrowly obovate, margins remotely shallowly dentate to subentire; bracts spreading, flat. |
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Flowers | usually 1 opening per day near sunset; buds erect, with free tips erect and appressed or slightly spreading, 0.5–2 mm; floral tube 15–35 mm; sepals3–11 mm; petals yellow, very broadly obovate or obcordate, 4.5–16 mm; filaments 4–11 mm, anthers 2–5.5 mm, pollen ca. 50% fertile; style 23–45 mm, stigma surrounded by anthers at anthesis. |
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Capsules | cylindrical, sometimes slightly enlarged toward apex, 15–45 × 2–3 mm. |
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Seeds | usually ellipsoid to broadly ellipsoid, rarely subglobose, 1–2 × 0.5–0.9 mm. |
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2n | = 14. |
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Calylophus tubicula |
Oenothera humifusa |
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Phenology | Flowering Apr–Nov. | |
Habitat | Dunes, open sandy places along or near Atlantic coast. | |
Elevation | 0–10 m. (0–0 ft.) | |
Distribution |
sw United States; sc United States; n Mexico |
AL; DE; FL; GA; LA; MD; MS; NC; NJ; PA; SC; VA; Dunes; open sandy places along or near Atlantic coast; West Indies (Cuba); Bermuda
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Discussion | Subspecies 2 (1 in the flora). H. F. Towner (1977) found that Oenothera tubicula is self-incompatible and diurnal with opening times just prior to sunrise. It occurs primarily on limestone soil in arid lowlands, but occasionally in montane areas, from Guadalupe County, New Mexico, south to western Texas, northeast to Howard County, Texas, and south to northern Zacatecas, south-central Nuevo León, and southwestern Tamaulipas, 600–1800 m. Subspecies strigulosa (Towner) W. L. Wagner & Hoch is known only from rocky, open sites and canyons in relatively montane areas, sometimes in pine forests in southernmost Coahuila, south-central Nuevo León, and southeastern Tamaulipas, from 1500 to 2300 m. It differs in being strigillose on the ovary and distally on stems, leaves linear to narrowly lanceolate, and the petals fading red or purple. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera humifusa is a PTH species and forms a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous (W. Dietrich and W. L. Wagner 1988). The inland collection from Iredell County, North Carolina, presumably represents an introduction. There are two geographically separated morphological forms of O. humifusa. Plants of one form are somewhat decumbent, with subentire cauline leaves and bracts; this form occurs in the southern part of the range. The other form is more upright, with more deeply divided leaves; it occurs from North Carolina northward. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | Calylophus tubiculus, Galpinsia tubicula, O. hartwegii var. tubicula | O. niveifolia, O. sinuata var. humifusa, Raimannia humifusa |
Name authority | A. Gray: Smithsonian Contr. Knowl. 3(5): 71. (1852) | Nuttall: Gen. N. Amer. Pl. 1: 245. (1818) |
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