Oenothera rosea |
Oenothera cespitosa |
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pink evening primrose, rose evening-primrose |
fragrant evening-primrose, tuft evening-primrose |
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Habit | Herbs perennial, caulescent, strigillose and often also sparsely hirsute; from slender taproot. | Herbs perennial, acaulescent or caulescent, usually hirsute or villous, usually also glandular puberulent, or exclusively strigillose, rarely glabrous; from stout taproot, sometimes lateral roots producing adventitious shoots. | ||||||||||||||||||||||||||||
Stems | 1–several, ascending to decumbent, 7–65 cm. |
(when present), usually ascending or decumbent, unbranched or branched from near base, 0–40 cm. |
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Leaves | in a basal rosette and cauline, basal 1–6 × 0.3–2 cm, blade narrowly elliptic to elliptic or ovate, margins subentire, weakly serrulate, or sinuate-pinnatifid; cauline 1–6 × 0.3–2 cm, blade narrowly elliptic to narrowly ovate, margins subentire or weakly serrulate, proximal ones sinuate-pinnatifid. |
1.7–26(–36) × (0.3–)0.5–4.5(–6.5) cm; petiole (0.2–)1.7–11(–14) cm; blade usually oblanceolate to rhombic or spatulate, rarely elliptic, obovate, lanceolate, or linear-oblanceolate, margins irregularly sinuate-dentate, serrate, pinnatifid, lobed, or subentire, apex usually acute to rounded, rarely acuminate. |
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Inflorescences | erect. |
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Flowers | 1–3 opening per day near sunrise; buds with free tips 0.1–1 mm; floral tube 4–8 mm; sepals 6–12 mm; petals rose purple, fading darker, 4–12 mm; filaments 4–6 mm, anthers 2–3.5 mm, pollen 35–65% fertile; style 7–13.5 mm, stigma surrounded by anthers at anthesis. |
1–4(–6) per stem opening per day near sunset, with moderate to strong sweet scent with a rubbery background scent; buds usually erect, rarely recurved (during early development); floral tube (20–)40–140(–165) mm; sepals (15–)18–45(–54) mm; petals white, fading rose or rose pink to dark or deep rose purple, or pink to pale or light rose, or lavender, obovate or obcordate, (16–)20–50(–60) mm; filaments (6–)10–30(–35) mm, anthers (6–)9–17(–20) mm; style (45–)60–180(–185) mm, stigma exserted beyond anthers at anthesis. |
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Capsules | narrowly obovoid, 4–12 × 2–4 mm, apex attenuate to a sterile beak, proximal stipe 5–20 mm, gradually tapering to base, valve midrib prominent in distal part; sessile. |
straight, curved, falcate, or sigmoid, usually cylindrical to lanceoloid or ellipsoid, sometimes ovoid, usually obtusely 4-angled, (10–)13–50(–68) × 4–9 mm, tapering to a sterile beak 6–8 mm, valve margins with rows of distinct tubercles to sinuate or nearly smooth ridges, dehiscent 1/3–7/8 their length; pedicel (0–)1–40(–55) mm. |
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Seeds | narrowly obovoid, 0.5–0.9 × 0.3–0.5 mm. |
numerous in 1 or 2 rows per locule, usually obovoid, oblong, or triangular, rarely suborbicular, 2.1–3.9 × 1–2.6 mm, embryo 1/5–2/3 of seed volume, surface papillose, reticulate or rarely irregularly roughened; seed collar sealed by a thin membrane, this flat or depressed into raphial cavity, when depressed often splitting, becoming separated from seed collar. |
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2n | = 14. |
= 14, 28. |
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Oenothera rosea |
Oenothera cespitosa |
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Phenology | Flowering Mar–Sep. | |||||||||||||||||||||||||||||
Habitat | tropical areas.. | |||||||||||||||||||||||||||||
Elevation | 10–600 m. (0–2000 ft.) | |||||||||||||||||||||||||||||
Distribution |
AZ; CA; TX; Mexico; Central America; West Indies; tropical areas [Introduced in South America (Argentina), Europe, Asia, s Africa, Atlantic Islands (Azores, Canary Islands)]
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w North America; nw Mexico
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Discussion | Oenothera rosea is a PTH species, forming a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous. In the flora area, it is known from Cochise, Pima, and Santa Cruz counties in Arizona, Alameda, Los Angeles, Riverside, San Bernardino, San Diego, San Francisco, and Santa Barbara counties in California (primarily in urban areas), and from southern Texas. It is clearly of North American origin, since all of its close relatives are confined to North America, and has spread south along the Andes. It occurs at 500–3700 m in South America but generally at lower elevations in most areas. The name Hartmannia affinis Spach is illegitimate, being based on Oenothera virgata; H. gauroides Spach is also illegitimate, being based on O. rosea; O. purpurea Lamarck is a later homonym; these three names pertain here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 5 (5 in the flora). Oenothera cespitosa occurs in a wide array of habitats, from grassland, desert scrub, pinyon-juniper woodland, or Arizona chaparral to montane conifer forests, rarely at timberline, at elevations from (450–)800–3370 m. Oenothera cespitosa is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005). Pachylophus nuttallii Spach is an illegitimate name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||||||||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Hartmannia | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Pachylophus | ||||||||||||||||||||||||||||
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Synonyms | Gaura epilobia, Godetia heuckii, Hartmannia rosea, H. rosea var. parvifolia, H. virgata, O. psycrophila, O. rosea var. parvifolia, O. rubra, O. virgata, Xylopleurum roseum | Pachylophus cespitosus | ||||||||||||||||||||||||||||
Name authority | L’Héritier ex Aiton: Hort. Kew. 2: 3. (1789) | Nuttall: Cat. Pl. Upper Louisiana, no. 53. (1813) | ||||||||||||||||||||||||||||
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