Oenothera rosea |
Oenothera californica |
|||||||||
---|---|---|---|---|---|---|---|---|---|---|
pink evening primrose, rose evening-primrose |
California evening primrose |
|||||||||
Habit | Herbs perennial, caulescent, strigillose and often also sparsely hirsute; from slender taproot. | Herbs perennial, densely strigillose, sometimes also villous, or glabrous; from a taproot, lateral roots producing adventitious shoots, or rarely with fleshy underground horizontal rootstock (subsp. eurekensis). | ||||||||
Stems | 1–several, ascending to decumbent, 7–65 cm. |
ascending or decumbent, usually branched from near base, sometimes new rosettes forming at branch apex when buried in drifting sand, 10–60 cm. |
||||||||
Leaves | in a basal rosette and cauline, basal 1–6 × 0.3–2 cm, blade narrowly elliptic to elliptic or ovate, margins subentire, weakly serrulate, or sinuate-pinnatifid; cauline 1–6 × 0.3–2 cm, blade narrowly elliptic to narrowly ovate, margins subentire or weakly serrulate, proximal ones sinuate-pinnatifid. |
in a basal rosette and cauline, rosette sometimes weakly developed or absent, at least during flowering, 1–13 × 0.5–2 cm; petiole 0–2(–4.5) cm; blade oblong to oblanceolate or spatulate, sometimes rhombic-ovate, margins entire or weakly to conspicuously dentate or pinnatifid. |
||||||||
Inflorescences | erect. |
|||||||||
Flowers | 1–3 opening per day near sunrise; buds with free tips 0.1–1 mm; floral tube 4–8 mm; sepals 6–12 mm; petals rose purple, fading darker, 4–12 mm; filaments 4–6 mm, anthers 2–3.5 mm, pollen 35–65% fertile; style 7–13.5 mm, stigma surrounded by anthers at anthesis. |
1–several opening per day near sunset; buds nodding, weakly quadrangular, with free tips 0–0.8 mm; floral tube 20–40 mm; sepals 15–30 mm, not spotted; petals white, fading pink to deep pink, broadly obcordate, 15–35(–40) mm; filaments 10–17 mm, anthers 5–10 mm; style 30–60 mm, stigma exserted beyond anthers at anthesis. |
||||||||
Capsules | narrowly obovoid, 4–12 × 2–4 mm, apex attenuate to a sterile beak, proximal stipe 5–20 mm, gradually tapering to base, valve midrib prominent in distal part; sessile. |
spreading to ascending, woody in age, often curved upward, cylindrical, obtusely 4-angled, tapering slightly from base to apex, 20–80 × 2–3.5 mm; sessile. |
||||||||
Seeds | narrowly obovoid, 0.5–0.9 × 0.3–0.5 mm. |
numerous, in 1 row per locule, olive-brown or yellowish brown to black, sometimes with minute purple dots, obovoid, 1–2.5 mm. |
||||||||
2n | = 14. |
|||||||||
Oenothera rosea |
Oenothera californica |
|||||||||
Phenology | Flowering Mar–Sep. | |||||||||
Habitat | tropical areas.. | |||||||||
Elevation | 10–600 m. (0–2000 ft.) | |||||||||
Distribution |
AZ; CA; TX; Mexico; Central America; West Indies; tropical areas [Introduced in South America (Argentina), Europe, Asia, s Africa, Atlantic Islands (Azores, Canary Islands)]
|
w United States; nw Mexico
|
||||||||
Discussion | Oenothera rosea is a PTH species, forming a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous. In the flora area, it is known from Cochise, Pima, and Santa Cruz counties in Arizona, Alameda, Los Angeles, Riverside, San Bernardino, San Diego, San Francisco, and Santa Barbara counties in California (primarily in urban areas), and from southern Texas. It is clearly of North American origin, since all of its close relatives are confined to North America, and has spread south along the Andes. It occurs at 500–3700 m in South America but generally at lower elevations in most areas. The name Hartmannia affinis Spach is illegitimate, being based on Oenothera virgata; H. gauroides Spach is also illegitimate, being based on O. rosea; O. purpurea Lamarck is a later homonym; these three names pertain here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (3 in the flora). Most populations of Oenothera californica are self-incompatible (W. L. Wagner et al. 2007; K. E. Theiss et al. 2010), but some populations of subsp. californica are self-compatible. All chromosome counts indicate that subspp. avita and eurekensis are diploid (2n = 14) and those of subsp. californica are tetraploid (2n = 28). Oenothera californica is polymorphic with subspp. avita and californica being very similar, and differing primarily in ecology, distribution, and relatively minor differences in leaf morphology and ploidy level, while the sand dune-restricted subsp. eurekensis is more distinctive in both morphology and habitat. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||
Key |
|
|||||||||
Source | FNA vol. 10. | FNA vol. 10. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Gaura epilobia, Godetia heuckii, Hartmannia rosea, H. rosea var. parvifolia, H. virgata, O. psycrophila, O. rosea var. parvifolia, O. rubra, O. virgata, Xylopleurum roseum | O. albicaulis var. californica, Anogra californica, O. pallida var. californica | ||||||||
Name authority | L’Héritier ex Aiton: Hort. Kew. 2: 3. (1789) | (S. Watson) S. Watson in W. H. Brewer et al.: Bot. California 1: 223. (1876) | ||||||||
Web links |