Oenothera pubescens |
Oenothera heterophylla |
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pubescent evening primrose, silky evening primrose, South American evening-primrose |
variableleaf evening primrose |
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Habit | Herbs annual or biennial, densely to sparsely strigillose, sometimes also villous and glandular puberulent distally; from a taproot. | Herbs annual or short-lived perennial, sparsely to densely strigillose, inflorescence sometimes also sparsely glandular puberulent, villous, or sparsely hirsute with spreading, pustulate-based hairs, or sometimes glabrate. | ||||
Stems | unbranched or with branched central stem and ascending to decumbent lateral branches arising from rosette, 5–50(–80) cm. |
unbranched or branched mainly in distal part, 25–70 cm. |
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Leaves | in a basal rosette and cauline, basal 5–14 × 0.5–2.5 cm, cauline 2–8 × 0.5–2.5 cm; blade narrowly oblanceolate to narrowly elliptic or narrowly oblong, margins usually dentate to deeply lobed; bracts spreading, flat. |
in a basal rosette and cauline, basal 7–15 × 1–2.5 cm, cauline 3–13 ×0.4–2.3 cm; blade narrowly oblanceolate to oblanceolate, gradually narrowly lanceolate to lanceolate or elliptic distally, margins deeply lobed to remotely dentate or subentire; bracts longer than capsule they subtend, 1–3 cm. |
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Inflorescences | dense, often with several lateral branches, mature buds usually overtopping spike apex. |
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Flowers | usually 1 opening per day near sunset; buds with free tips erect, 0.1–1 mm; floral tube erect, becoming recurved and nodding, then erect again just before anthesis, 15–50 mm; sepals 5–25 mm; petals yellow, fading reddish orange, broadly obovate to obcordate, 5–25(–35) mm; filaments 6–18 mm, anthers (2–)3–9 mm, pollen ca. 50% fertile; style 20–60 mm, stigma surrounded by or slightly exserted beyond anthers at anthesis. |
2–several per spike opening per day near sunset; buds erect, with free tips erect or spreading, 1–6 mm; floral tube nearly straight, 25–47 mm; sepals 15–30 mm; petals yellow, broadly elliptic to nearly rhombic, 18–35 mm; filaments 15–30 mm, anthers 3–8 mm, pollen 85–100% fertile; style 45–75 mm, stigma usually exserted beyond anthers at anthesis. |
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Capsules | cylindrical, sometimes slightly enlarged distally, 20–45 ×2–4 mm. |
lanceoloid, 13–25 × 2.5–4 mm. |
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Seeds | brown, sometimes dark-flecked, 0.9–1.5 × 0.6–1 mm. |
brown, often flecked with darker spots, ellipsoid to broadly ellipsoid, 1.1–1.8 × 0.4–0.8 mm. |
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2n | = 14. |
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Oenothera pubescens |
Oenothera heterophylla |
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Phenology | Flowering (Feb–)Apr–Sep(–Oct). | |||||
Habitat | Open sites in montane habitats. | |||||
Elevation | (1300–)1500–2500(–3100) m. ((4300–)4900–8200(–10200) ft.) | |||||
Distribution |
AZ; NM; TX; Mexico; West Indies; Central America (Guatemala); South America (Colombia, Ecuador, Peru)
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s United States |
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Discussion | Oenothera pubescens is a PTH species and forms a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous (W. Dietrich and W. L. Wagner 1988). Oenothera pubescens has been collected once in California in 1884 (Newberry Springs, San Bernardino County), where it was temporarily introduced or a natural occurrence that was extirpated. Collections from west Texas (Brewster, Jeff Davis, and Presidio counties) have been made since 1990 and a few others collected earlier were misidentified as O. laciniata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Nutantigemma | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Candela | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Anogra amplexicaulis, O. amplexicaulis, O. laciniata subsp. pubescens, Var. o. var. o., O. nyctaginiifolia, O. stuebelii, Raimannia colimae, R. confusa | Raimannia heterophylla | ||||
Name authority | Willdenow ex Sprengel: Syst. Veg. 2: 229. (1825) | Spach: Nouv. Ann. Mus. Hist. Nat. 4: 348. (1836) | ||||
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