Oenothera psammophila
Oenothera triloba
St. Anthony dunes evening primrose
stemless evening-primrose
decumbent, usually branched, 10–30 cm, becoming woody and buried in sand.
(when present) ascending, 1–several, densely leafy, 0–20 cm.
(6–)8–9(–14.2) × (0.7–)1.5–2(–3.1) cm;
petiole 3–9 cm;
blade narrowly oblanceolate to oblanceolate, margins usually subentire or repand or remotely dentate, rarely serrate, apex acute.
in a basal rosette, sometimes also cauline, (2.5–)6–25(–32) × (0.6–)1.5–4(–5) cm, thin;
petiole (0.5–)1–8 cm;
blade oblanceolate to elliptic, margins irregularly pinnatifid, sometimes subentire, apex acute to obtuse or rounded.
1–3 per stem opening per day near sunset, with sweet scent;
buds erect;
floral tube 42–60 mm;
sepals 22–28 mm;
petals white, fading rose pink to rose, 23–40 mm;
filaments 17–19 mm, anthers 13–16 mm;
style 60–72(–88) mm, stigma exserted beyond anthers at anthesis.
1–4 opening per day near sunset, without noticeable scent;
buds with subequal free tips 2–7 mm;
floral tube (20–)28–95(–138) mm;
sepals (6–)10–30(–35) mm;
petals pale yellow, fading pale orange, drying lavender, (10–)12–30(–38) mm;
filaments (5–)8–15(–18) mm, anthers (3.5–)4–11 mm;
style (3.4–)4.2–11.5(–16.3) mm, stigma usually surrounded by anthers, sometimes (especially in some Texas populations) exserted beyond anthers.
somewhat curved and often somewhat twisted, lanceoloid-cylindrical, nearly cylindrical, (20–)30–47 × 7–8 mm, gradually tapering to apex, 6–8 mm, dehiscent nearly throughout their length, valve margins with a conspicuous, irregular, wavy ridge;
pedicel 1–5 mm.
woody in age, rhombic-obovoid, winged, wings broadly triangular, 5–10 mm wide, often terminating in a hooked tooth, (10–)15–25(–28) × 4–8 mm (excluding wings), valve surface reticulate, dehiscent 1/8–1/3 their length.
numerous, in 2 distinct rows per locule, narrowly obovoid, 2.5–3 × 1.2–1.4 mm, embryo 7/8 of seed volume, surface finely reticulate;
seed collar with a broad membrane sealing cavity, margin entire.
asymmetrically cuneiform, (2.1–)2.5–3(–3.3) mm.
= 14.
= 14.
Oenothera psammophila
Oenothera triloba
Oenothera psammophila is known only from the dune area north and west of St. Anthony, Fremont County. It is unique in Oenothera because of the exudate produced on the leaves to which sand particles adhere, forming a sand sheath, presumably for protection from the constantly blowing sand particles. It is self-incompatible (W. L. Wagner et al. 1985; Wagner 2005).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Oenotheratriloba is primarily a species of the high plains from eastern Socorro County, New Mexico, east through all but eastern Texas, Oklahoma, to southern Kansas, east of Meade and Pawnee counties and south of Douglas and Saline counties. It becomes more sporadic eastward into Missouri south of the Missouri River, northwestern and north-central Arkansas, central and eastern Tennessee, northern Alabama, and Logan and Warren counties, Kentucky; also known from disjunct sites in northern Mexico from Nuevo León, Chihuahua, and Baja California, Mexico; and, introduced in Illinois, Indiana, Ohio, Kentucky (Campbell and Fayette counties), Pennsylvania, Virginia, Maryland, and the District of Columbia. Areas where it was introduced are represented by old collections; no current information indicates their continued presence in any of these areas. It was recently collected in Baca County, Colorado.
Capsules of dead plants sometimes form pineconelike clusters of ten to 100 or more capsules.
The illegitimate names Lavauxia nuttalliana Spach and L. triloba (Nuttall) Spach var. watsonii Britton pertain here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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