Oenothera primiveris |
Oenothera canescens |
|
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desert evening primrose, yellow desert evening primrose |
beakpod evening primrose, spotted evening-primrose |
|
Habit | Herbs winter-annual, caulescent to short-caulescent, long-hirsute, hairs often with reddish purple pustulate bases, especially proximally, also moderately strigillose, and glandular puberulent distally, often on leaves; from a weakly fleshy taproot. | Herbs low, forming clumps 10–50 cm diam., densely strigillose throughout; from a taproot, lateral roots producing adventitious shoots. |
Stems | (when present) unbranched and erect or, sometimes, few branches from near base, in robust plants stems and caudex hollow and greatly enlarged, especially toward base, densely leafy, 5–35 cm. |
many-branched from base, leafy, (10–)15–25(–38) cm. |
Leaves | in a basal rosette, sometimes also cauline, (1.4–)6–15(–28) × (0.2–)1–3.5(–5.6) cm; petiole (0.9–)3.5–8(–14) cm; blade oblanceolate to linear-oblanceolate, pinnatifid or 2-pinnatifid to shallowly pinnately lobed, margins sinuate-dentate or subentire, apex obtuse. |
cauline, (0.3–)0.6–1.5(–2.5) × (0.05–)0.15–0.4(–0.6) cm, fascicles of small leaves 0.2–0.6 cm often present in non-flowering axils; petiole 0–0.1 cm; blade lanceolate to linear, base cuneate, apex acute. |
Flowers | usually 1–4, rarely more, opening per day, 1–2 hours before sunset; sepals (7–)12–25(–30) mm; petals yellow, fading reddish orange to purple, obcordate to obovate, (6–)13–35(–40) mm; filaments 6–16 mm, anthers 3–10 mm; style (32–)40–90(–100) mm, stigma exserted beyond anthers or surrounded by them. |
several opening per day near sunset; buds usually without free tips, rarely free tips 0.2–0.3 mm; sepals (7–)8–12 mm; petals pink, rarely white, streaked or flecked with red, fading bright purple, (8–)10–17 mm; filaments 6–8 mm, anthers often with red longitudinal stripe, 3–6 mm; style (16–)22–27 mm, stigma exserted beyond anthers at anthesis. |
Capsules | woody in age, sigmoid or curved to nearly straight, lanceoloid to ovoid, 4-angled, 10–45(–60) × 4–8 mm, beak 4–15 mm, dehiscent 1/4–2/3 their length; sessile. |
woody, ovoid, narrowly winged, wings 0.8–1.5 mm wide, (7–)9–12(–14) × 2–4 mm (excluding wings), abruptly constricted to a conspicuous, sterile beak, (2–)3–4.5 mm, indehiscent; sessile. |
Seeds | usually numerous, in 2 rows per locule, obovoid to oblanceoloid, 3–3.5 × 1–1.4 mm, surface thickened above raphe and at distal end into U-shaped structure. |
asymmetrically cuneiform or oblanceoloid, 1.2–1.5 × 0.4–0.5 mm. |
2n | = 14. |
= 14. |
Oenothera primiveris |
Oenothera canescens |
|
Phenology | Flowering Feb–May(–Jun). | Flowering May–Aug. |
Habitat | Sandy soil on flats, low hills and margins of sand dunes, along arroyos, roadsides, in desert scrub, grasslands and oak-grasslands. | Prairie depressions, playas, margins of ditches, temporary wet areas. |
Elevation | 30–1600 m. (100–5200 ft.) | (400–)700–1800 m. ((1300–)2300–5900 ft.) |
Distribution |
AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Sonora)
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CO; KS; NE; NM; OK; TX; WY
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Discussion | Oenothera primiveris has a complex variation pattern (W. L. Wagner 2005). In the western part of the range from southeastern California across southern Nevada to southern Utah counties of Emery, Kane, and Washington, and northwestern Mohave County, Arizona, plants generally have a gray appearance, with dense pubescence and larger flowers with widespread self-compatibility, but with scattered populations retaining self-incompatibility. Populations from southof the Mogollon Plateau to southern New Mexico, western Texas, Chihuahua, Sonora, and Baja California, Mexico, are greener in appearance with smaller to much smaller flowers, and are all self-compatible with occasional outcrossing or complete autogamy. The transitions between these two extremes are so extensive and more or less gradual that it is not possible to subdivide into two subspecies as has been done previously (Wagner). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera canescens is restricted to prairie depressions, playas, ditch margins, and other places of temporary water in the High Plains of the western United States from Goshen County, Wyoming, southeast to Hayes County, Nebraska, south through eastern Colorado, the eastern tier of counties in New Mexico, western Kansas, and to Garza and Dawson counties in the Texas Panhandle; also disjunct populations from Chautauqua, Sedgwick, and Stafford counties, Kansas. The illegitimate names Gaurella guttulata (Geyer ex Hooker) Small, G. canescens (Torrey & Frémont) Cockerell, and Gauropsis guttulata (Geyer ex Hooker) Cockerell pertain here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Eremia | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gauropsis |
Sibling taxa | ||
Synonyms | Lavauxia lobata, L. primiveris, O. bufonis, O. cespitosa var. primiveris, O. johnsonii, O. primiveris subsp. bufonis, O. primiveris var. bufonis, O. primiveris subsp. caulescens, O. primiveris var. caulescens, Pachylophus johnsonii | Gaurella canescens, Megapterium canescens, O. guttulata |
Name authority | A. Gray: Smithsonian Contr. Knowl. 5(6): 58. (1853) | Torrey & Frémont in J. C. Fremont: Rep. Exped. Rocky Mts., 315. (1845) |
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