Oenothera platanorum
Oenothera macrocarpa
Fort Huachuca evening primrose
bigfruit evening primrose, Missouri evening primrose, Ozark sundrop
1–several, ascending, 5–60 cm.
moderately leafy, (1–)4–40(–60) cm.
in a basal rosette and cauline, basal 2–7 × 0.3–1.4 cm, blade narrowly elliptic to narrowly ovate, margins weakly serrulate to sinuate-pinnatifid;
cauline 1.2–6 × 0.3–1 cm, blade narrowly elliptic to elliptic or ovate, proximal ones sinuate-pinnatifid, margins subentire or weakly serrulate.
cauline, (2.8–)3.7–12.5(–17) × (0.1–)0.4–3(–4.5) cm;
petiole (0.4–)1–4(–6) cm;
blade linear, lanceolate-elliptic, elliptic to oblanceolate or suborbiculate, margins entire or conspicuously or inconspicuously denticulate or serrulate, sometimes undulate, apex usually acute, sometimes obtuse or retuse (subsp. incana).
erect.
1–3 opening per day near sunrise;
buds with free tips 0–0.1 mm;
floral tube 9–14 mm;
sepals 7.5–13 mm;
petals rose purple, fading darker, 8–15 mm;
filaments 4–9 mm, anthers 2.5–4 mm, pollen 85–100% fertile;
style 12–19 mm, stigma surrounded by anthers at anthesis.
usually 1 or 2, rarely more, opening per day near sunset, fading next morning, sometimes (subspp.
clavate or narrowly obovoid, 9–14 × 3–4 mm, apex attenuate to a sterile beak, valve midrib prominent in distal part, proximal stipe 4–15 mm, gradually tapering to base;
sessile.
papery in age, narrowly ellipsoid to lanceoloid, sometimes twisted (subsp. fremontii), winged, wings (2–)10–28(–34) mm wide, body (13–)25–70(–115) × 2–9 mm, dehiscent 1/4–1/3 their length;
pedicel 1–12(–25) mm.
narrowly obovoid, 0.7–0.9 × 0.3–0.5 mm.
numerous, rarely as few as 8, in 1 row per locule, obovoid, (2–)3–5 × 1–2.3 mm.
and oklahomensis) lasting for 2 days, weakly scented;
buds with unequal free tips 1–11(–15) mm;
floral tube (21–)35–140(–160) mm;
sepals (20–)25–65(–75) mm;
petals bright yellow, fading orange, reddish orange or mostly unchanged, obovate to very broadly obovate, (17–)25–65(–68) mm, usually with terminal notch and/or tooth, margin sometimes erose;
filaments 13–40(–44) mm, anthers 10–24(–25) mm;
style (45–)55–192 mm, stigma usually exserted beyond anthers at anthesis.
= 14.
Oenothera platanorum
Oenothera macrocarpa
Oenothera platanorum is known only from the southeastern counties of Cochise, Pinal, and Santa Cruz in Arizona. It was recently collected in Sonora, Mexico. The species is very similar to both O. texensis, from which it differs in its smaller flowers, and the widespread O. rosea, from which it differs in the somewhat larger flowers and in forming seven bivalents in meiosis and fully fertile pollen, whereas O. rosea is a PTH species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 5 (4 in the flora).
Oenotheramacrocarpa is variable and has differentiated extensively in the Great Plains region. Each of the five distinctive subspecies occupies a different geographical and ecological situation. Only subsp. mexicana W. L. Wagner from Coahuila, Mexico, occurs outside of the flora area. In general, the subspecies are sharply distinct and each is characterized by a number of features, including pubescence, leaf features, flower and floral tube size, and size and morphology of the capsules and seeds. The five entities are treated as subspecies primarily because of their complete interfertility and extensive intergradation in any area of marginal contact. Intermediates are known between subsp. macrocarpa and subspp. fremontii and oklahomensis and between subspp. incana and oklahomensis. There is also some evidence that suggests past hybridization between subspp. fremontii and incana although there is no present contact between them. All subspecies are self-incompatible.
Oenothera alata Nuttall (1818) is an illegitimate name based on O. macrocarpa and pertains here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Herbs glabrous. | → 2 |
2. Leaf blades usually very broadly elliptic to suborbiculate, rarely oblanceolate or elliptic, 2–4.3 cm wide, margins usually entire, sometimes inconspicuously denticulate, usually flat, sometimes undulate. | subsp. incana |
2. Leaf blades usually elliptic to lanceolate-elliptic, sometimes linear, (0.3–)0.8–2(–3) cm wide, margins usually conspicuously denticulate to serrulate, rarely subentire, usually undulate. | subsp. oklahomensis |
1. Herbs strigillose. | → 3 |
3. Bud free tips 1–2(–5) mm; petals (17–)25–33(–37) mm; floral tubes (21–)35–65(–80) mm; capsules often twisted, 13–30(–65) mm, wings 2–5(–9) mm wide. | subsp. fremontii |
3. Bud free tips (4–)5–11(–12) mm; petals (25–)31–65(–68) mm; floral tubes (50–)70–140(–160) mm; capsules not twisted, (28–)30–70(–115) mm, wings 10–28(–34) mm wide. | → 4 |
4. Leaves green or younger ones grayish green, moderately strigillose, rarely more densely strigillose, blades often lanceolate-elliptic to broadly elliptic, sometimes linear or lanceolate, 0.4–2.3(–3) cm wide; capsule wings (14–)18–28(–34) mm wide. | subsp. macrocarpa |
4. Leaves gray, densely strigillose, blades very broadly elliptic to suborbiculate, rarely oblanceolate or elliptic, 2–4.3 cm wide; capsule wings 10–15(–24) mm wide. | subsp. incana |
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