Oenothera pilosella |
Oenothera engelmannii |
|
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meadow evening-primrose |
Engelmann's evening-primrose |
|
Habit | Herbs perennial, usually densely to sparsely hirsute, rarely glabrous; from a thickened base, rhizomatous. | Herbs winter-annual, conspicuously villous throughout, also strigillose on leaves and distal parts; from a taproot. |
Stems | spreading or ascending, unbranched or few-branched distally, 20–80 cm. |
erect, unbranched or with few, spreading branches, 30–50(–80) cm. |
Leaves | in a basal rosette and cauline, basal 4–8 × 2–5 cm, petiole (0.5–)1–3(–4) cm, blade oblanceolate to ovate, margins entire; cauline 3–10(–13) × 1–2(–4) cm, petiole 0–0.5(–2) cm, blade lanceolate to ovate, abruptly narrowed to base, margins subentire or coarsely dentate. |
in a basal rosette and cauline, rosette weakly developed or absent, at least during flowering, (1–)2–6(–8) × 1–2(–3) cm; sessile; blade lanceolate to oblong lanceolate, proximal ones sometimes oblanceolate, margins coarsely repand-dentate or -pinnatifid. |
Inflorescences | erect, flowers in axils of distalmost few nodes. |
|
Flowers | opening near sunrise; buds with free tips 1–3 mm, spreading; floral tube 10–25 mm; sepals 10–20 mm; petals bright yellow, fading pale pink or pale yellow, 15–30 mm; filaments 7–15 mm, anthers 4–8 mm, pollen 85–100% fertile; style 10–20 mm, stigma exserted beyond anthers at anthesis. |
1–several opening per day near sunset; buds nodding, weakly quadrangular, without free tips; floral tube 20–30 mm; sepals 13–21 mm, not spotted or with scattered small, maroon spots; petals white, fading pink, broadly obovate or obcordate, 15–30 mm; filaments 14–16 mm, anthers 6–8 mm; style 40–50 mm, stigma exserted beyond anthers at anthesis. |
Capsules | usually oblong-clavate to oblong-ellipsoid or ellipsoid, 4-angled or weakly 4-winged, (5–)10–15(–28) × 2–4(–5) mm, stipe (1–)3–5(–9) mm; sessile. |
widely spreading, woody in age, straight or slightly curved, cylindrical, obtusely 4-angled, especially toward base, tapering gradually from base to apex, 30–60 × 2–3 mm; sessile. |
Seeds | 1 × 0.5 mm. |
numerous, in 1 row per locule, brown, narrowly obovoid, 1–1.5 mm. |
2n | = 56. |
= 14. |
Oenothera pilosella |
Oenothera engelmannii |
|
Phenology | Flowering May–Jul(–Aug). | Flowering Apr–Aug(–Sep). |
Habitat | Open fields, edge of woods, marshes and bottomland prairies, open disturbed sites, ditches, old fields, railroads, roadsides. | Sandy prairies, dunes, disturbed areas, roadsides. |
Elevation | 100–600 m. (300–2000 ft.) | 500–1300 m. (1600–4300 ft.) |
Distribution |
AL; AR; CT; IA; IL; IN; KY; LA; MA; ME; MI; MO; MS; NH; NY; OH; OK; PA; RI; VA; VT; WI; WV; ON
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CO; KS; NM; OK; TX
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Discussion | Oenothera pilosella is widespread in cultivation in gardens and frequently escapes and becomes naturalized; the northern and eastern natural limits of O. pilosella are not clear. According to G. B. Straley (1977) the natural limits are from Wayne County, West Virginia, along the Ohio River and Erie County, New York, for the eastern limits, and Tuscola County, Michigan, and Manitowoc County, Wisconsin, for the northern limits. K. N. Krakos (2014), based on new field studies and phylogenetic data, found that O. pilosella does not form a monophyletic group with plants previously treated by Straley as O. pilosella subsp. sessilis in molecular analyses, and thus is here reinstated as the distinct species O. sessilis. Straley determined that O. pilosella is self-incompatible and an octoploid, one of the few in the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera engelmannii is self-incompatible (W. L. Wagner et al. 2007; K. E. Theiss et al. 2010). It has a relatively narrow distribution in sandy areas of eastern New Mexico and western Texas, extending to southeastern Colorado, western Oklahoma, and southwestern Kansas. The flower size seems to vary, with larger flowers in eastern New Mexico and considerably smaller flowers in the eastern part of its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Kneiffia | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Anogra |
Sibling taxa | ||
Synonyms | Kneiffia fruticosa var. pilosella, K. pilosella, K. pratensis, K. sumstinei, O. fruticosa var. hirsuta, O. fruticosa var. pilosella, O. pratensis | Anogra pallida var. engelmannii, A. engelmannii |
Name authority | Rafinesque: Ann. Nat. 1: 15. (1820) | (Small) Munz: Amer. J. Bot. 18: 316. (1931) — (as engelmanni) |
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