Oenothera parviflora |
Oenothera heterophylla |
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northern evening-primrose, small-flower evening-primrose |
variableleaf evening primrose |
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Habit | Herbs biennial, sparsely strigillose, glandular puberulent, and villous with pustulate or non-pustulate hairs, sometimes predominately strigillose proximally or predominately villous with pustulate or non-pustulate hairs distally, glabrous, or some mixture of strigillose, glandular puberulent, or sparsely villous distally, sometimes appearing glabrous to the naked eye. | Herbs annual or short-lived perennial, sparsely to densely strigillose, inflorescence sometimes also sparsely glandular puberulent, villous, or sparsely hirsute with spreading, pustulate-based hairs, or sometimes glabrate. | ||||
Stems | erect, green or red on proximal parts or throughout, mostly branched from base or only in distal 1/2, 30–150 cm. |
unbranched or branched mainly in distal part, 25–70 cm. |
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Leaves | in a basal rosette and cauline, basal 10–30 × 1–4 cm, cauline 4–18 × 1–3 cm; blade usually bright green, sometimes pale green distally, white- or red-veined, narrowly oblanceolate, narrowly elliptic to lanceolate, or oblong, margins usually flat, rarely undulate, regularly dentate to remotely denticulate, sometimes teeth widely spaced; bracts persistent. |
in a basal rosette and cauline, basal 7–15 × 1–2.5 cm, cauline 3–13 ×0.4–2.3 cm; blade narrowly oblanceolate to oblanceolate, gradually narrowly lanceolate to lanceolate or elliptic distally, margins deeply lobed to remotely dentate or subentire; bracts longer than capsule they subtend, 1–3 cm. |
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Inflorescences | erect or ± curved, unbranched or with secondary branches just proximal to main one. |
dense, often with several lateral branches, mature buds usually overtopping spike apex. |
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Flowers | opening near sunset; buds erect, 3–5 mm diam., with free tips subterminal, spreading to erect, 0.5–5 mm; floral tube 22–40 mm; sepals green to yellowish green or flushed with red or dark red, sometimes only red-flecked, 7–17 mm; petals yellow to pale yellow, fading pale yellow to pale yellowish orange, very broadly obcordate, 8–15(–20) mm; filaments 7–13 mm, anthers 3.5–6 mm, pollen ca. 50% fertile; style 25–50 mm, stigma surrounded by anthers at anthesis. |
2–several per spike opening per day near sunset; buds erect, with free tips erect or spreading, 1–6 mm; floral tube nearly straight, 25–47 mm; sepals 15–30 mm; petals yellow, broadly elliptic to nearly rhombic, 18–35 mm; filaments 15–30 mm, anthers 3–8 mm, pollen 85–100% fertile; style 45–75 mm, stigma usually exserted beyond anthers at anthesis. |
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Capsules | erect or slightly spreading, usually greenish black when dry, narrowly lanceoloid to lanceoloid, 20–40 ×3.5–5 mm, free tips of valves 1–1.5 mm. |
lanceoloid, 13–25 × 2.5–4 mm. |
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Seeds | 1.1–1.8 × 0.5–1 mm. |
brown, often flecked with darker spots, ellipsoid to broadly ellipsoid, 1.1–1.8 × 0.4–0.8 mm. |
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2n | = 14. |
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Oenothera parviflora |
Oenothera heterophylla |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Open or disturbed, sandy or gravelly sites, roadsides, fallow fields, clearings, riverbanks, along water courses, salt marshes, coastal meadows. | |||||
Elevation | 0–1700 m. (0–5600 ft.) | |||||
Distribution |
CT; DC; DE; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; ND; NH; NJ; NY; OH; PA; RI; TN; VA; VT; WI; WV; BC; NB; NF; NS; ON; PE; QC [Introduced in Europe, Asia (ne China, Japan), s Africa, Pacific Islands (New Zealand), Australia]
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s United States |
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Discussion | Oenothera parviflora is a PTH species and forms a ring of 14 chromosomes in meiosis, and is self-compatible and autogamous (W. Dietrich et al. 1997). It has plastome IV and a BC genome composition. Onagra chrysantha Spach 1835, not Michaux 1803, is a superfluous name, as is Onagra chrysantha var. parviflora (Linnaeus) Spach, and both pertain here. O. biformiflora var. cruciata R. R. Gates is an invalid name that pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Oenothera | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Candela | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | O. ammophiloidesr. var. flecticaulis, O. ammophiloides var. parva, O. angustifolia, O. angustissima, O. angustissima var. quebecensis, O. apicaborta, O. atrovirens, O. biennis var. cruciata, O. biennis var. parviflora, O. biformiflora, O. cleistantha, O. comosa, O. cruciata, O. cruciata var. sabulonensis, O. deflexa, O. flecticaulis, O. hazeliae, O. hazeliae var. parviflora, O. hazeliae var. subterminalis, O. intermedia, O. laevigata, O. laevigata var. scitula, O. laevigata var. similis, O. novae-scotiae, O. novae-scotiae var. distantifolia, O. novae-scotiae var. intermedia, O. parva, O. parviflora subsp. angustissima, O. parviflora var. angustissima, O. robinsonii, O. rubricapitata, O. scitula, O. subterminalis, O. venosa, Onagra biennis var. cruciata, O. chrysantha var. cruciata, O. cruciata, O. parviflora | Raimannia heterophylla | ||||
Name authority | Linnaeus: Syst. Nat. ed. 10, 2: 998. (1759) — (as parviflor) | Spach: Nouv. Ann. Mus. Hist. Nat. 4: 348. (1836) | ||||
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