Oenothera pallida subsp. runcinata
Oenothera pallida
mountain evening primrose, pale evening-primrose, rockweed brush, white-stem evening primrose
usually branched throughout.
erect or ascending, single to several from base, unbranched or many-branched throughout, 10–50(–70) cm.
rosette not present at anthesis, 2–3.5(–5) × 0.4–1(–1.5) cm;
blade oblong to narrowly lanceolate, margins usually deeply sinuate-dentate to pinnatifid, rarely dentate only.
cauline, rosette usually weakly developed or absent, at least during flowering, sometimes well developed, 1–5(–7.8) × 0.3–1(–1.5) cm;
petiole 0–2(–4.5) cm;
blade lanceolate, oblong, linear-lanceolate, or ovate, margins subentire or remotely denticulate, deeply sinuate-dentate, or pinnatifid, sometimes repand.
buds with free tips 0–0.2 mm;
floral tube 15–30 mm;
sepals 10–25 mm;
petals 10–30 mm.
1–several opening per day near sunset;
buds nodding, weakly quadrangular, with free tips 0–2 mm;
floral tube 15–40 mm;
sepals 10–30 mm, not spotted;
petals white, fading pink to deep pink, broadly obovate or obcordate, (10–)15–25(–40) mm;
filaments 9–15 mm, anthers 3–10 mm;
style 25–55 mm, stigma exserted beyond anthers at anthesis.
spreading to reflexed, straight or curved, sometimes contorted.
spreading to reflexed, straight to curved or contorted, cylindrical, obtusely 4-angled, tapering slightly from base to apex, 15–60 × 1.5–2.5 mm;
sessile.
numerous, in 1 row per locule, brownish with dark spots or black, narrowly obovoid, 1.5–2.2 mm.
= 14.
Oenothera pallida subsp. runcinata
Oenothera pallida
Subspecies runcinata replaces subsp. pallida in the southern part of the species range. In New Mexico, it is most morphologically diverse, with plants with subentire leaves densely strigillose recognized in the past as subsp. gypsophila. The distribution and characteristics of these plants should be studied more in relation to other types of populations included in subsp. runcinata. Glabrous plants in the southern portion of the range of subsp. runcinata, especially in New Mexico, adjacent Texas, and Chihuahua, Mexico, have been recognized as var. brevifolia.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 4 (4 in the flora).
Oenothera pallida is a poorly understood species currently subdivided into four subspecies (W. L. Wagner et al. 2007) that differ largely in aspect, leaf division, capsule configuration, and pubescence. The variation pattern is rather complex with almost no diagnostic character uniformly distinguishing any one of the subspecies. Instead, each of the subspecies, which are mostly geographically separated although there is some level of overlap, have diagnostic suites of characters that maintain their linkage some of the time, but break down across the geographic area of each so that no single character uniquely identifies it. Each subspecies is characterized by leaf, pubescence, and, often, habit features. The issues with the integrity and intergradations of the subspecies are discussed below.
Oenothera pallida has been determined to be self-incompatible (W. L. Wagner et al. 2007), but K. E. Theiss et al. (2010) determined that although most populations of subsp. pallida are self-incompatible, one near Salt Lake City is self-compatible.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Herbs annual, sometimes perennial from a taproot, when perennial, sometimes with lateral roots producing adventitious shoots, strigillose throughout and villous distally, especially on flower parts. | subsp. trichocalyx |
1. Herbs perennial from a taproot and with lateral roots producing adventitious shoots, glabrous, strigillose, or sparsely villous. | → 2 |
2. Plants glabrous, sometimes strigillose, rarely sparsely villous; leaf blade margins usually subentire or remotely denticulate, rarely pinnatifid; capsules usually contorted to curved. | subsp. pallida |
2. Plants usually strigillose, rarely villous or glabrous; leaf blade margins shallowly sinuate-dentate or denticulate, or deeply sinuate-dentate to pinnatifid, rarely only dentate; capsules usually straight or curved, sometimes contorted. | → 3 |
3. Leaf blades (0.4–)0.7–1.5 cm wide, margins shallowly sinuate-dentate or denticulate. | subsp. latifolia |
3. Leaf blades 0.4–1(–1.5) cm wide, margins usually deeply sinuate-dentate to pinnatifid, rarely dentate only. | subsp. runcinata |
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