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pale evening-primrose

Habit Herbs perennial, usually glabrous, sometimes strigillose, rarely sparsely villous; from a taproot, lateral roots producing adventitious shoots. Herbs (annual or perennial), [shrubs].
Stems

usually branched throughout.

Leaves

rosette not present at anthesis, 2–6 × 0.3–0.8(–1) cm;

blade lanceolate to linear-lanceolate or oblong, margins usually subentireor remotely denticulate, rarely pinnatifid, usually repand.

alternate or basal;

stipules absent.

Flowers

buds with free tips 0.5–2 mm;

floral tube 20–35 mm;

sepals 12–18 mm;

petals 12–25 mm.

usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;

stamens 2 times as many, or rarely as many, as sepals;

pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).

Fruit

a dry capsule, usually dehiscent, sometimes indehiscent.

Capsules

spreading, contorted to curved.

Seeds

few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).

2n

= 14.

Oenothera pallida subsp. pallida

Onagraceae tribe Onagreae

Phenology Flowering May–Sep.
Habitat Sandy soil, dunes, disturbed areas, alkaline soil.
Elevation 1100–2000 m. (3600–6600 ft.)
Distribution
from FNA
AZ; CO; ID; NM; NV; OR; UT; WA; WY; BC
[BONAP county map]
North America; Mexico; Central America; South America; West Indies
Discussion

The distribution of subsp. pallida centers in the intermountain region from Oregon and Washington east of the mountains, adjacent southern British Columbia, south through southern Idaho, Wyoming, western half of Utah, southern Nevada, to northern Arizona, northwestern New Mexico, and adjacent southwestern Colorado. There are morphological intermediates with subspp. runcinata and trichocalyx. Densely strigillose plants occur within the range of subsp. pallida, especially near the St. Anthony Dunes in Idaho, and have been referred to as var. idahoensis.

Baumannia douglasiana Spach is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 13, species 265 (12 genera, 199 species in the flora).

Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Anogra > Oenothera pallida Onagraceae > subfam. Onagroideae
Sibling taxa
O. pallida subsp. latifolia, O. pallida subsp. runcinata, O. pallida subsp. trichocalyx
Subordinate taxa
Synonyms Anogra leptophylla, O. pallida var. idahoensis, O. pallida var. leptophylla
Name authority unknown Dumortier: Fl. Belg., 89. (1827)
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