Oenothera oakesiana |
Oenothera cinerea |
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Oakes' evening-primrose |
high-plains beeblossom, woolly beeblossom |
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Habit | Herbs biennial, densely silky-strigillose, at least proximally, also sparsely villous with long, appressed hairs, sometimes also villous with spreading, pustulate hairs and/or glandular puberulent distally. | Herbs suffrutescent, densely soft-villous, hairs mostly appressed, 2–3 mm, becoming less villous distally, also strigillose, rarely glandular puberulent or hispidulous, plant parts grayish green; from deep, twisted, woody rootstock. | ||||
Stems | erect to decumbent, green or flushed with red on proximal parts or throughout, unbranched or bushy and branched from base, with side branches arising obliquely or arcuately from rosette, 10–60 cm. |
erect, several-branched near ground, also branched proximal to inflorescences, 60–280 cm. |
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Leaves | in a basal rosette and cauline, basal 8–30 × 0.5–3 cm, cauline 3.5–20 ×0.5–2.7 cm; blade grayish green to dull green, very narrowly oblanceolate to narrowly oblanceolate or narrowly elliptic, margins flat, subentire or remotely dentate, teeth sometimes blunt, sometimes sinuate-dentate proximally; bracts persistent. |
in a basal rosette and cauline, 0.5–8 × 0.15–2 cm, sessile, blade narrowly lanceolate or oblanceolate to very narrowly elliptic or linear, margins usually subentire or shallowly sinuate-dentate, sometimes deeply sinuate-dentate, often undulate. |
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Inflorescences | usually recurved with ascending tip distally, rarely suberect, unbranched. |
slender. |
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Flowers | opening near sunset; buds erect, 3–5 mm diam., with free tips subterminal, spreading to erect, 2.5–4 mm; floral tube 15–40 mm; sepals green to yellow, flushed with red and dark red flecked or red-striped, 9–17 mm; petals yellow to pale yellow, fading yellowish white to pale yellowish orange, very broadly obcordate, 7–20 mm; filaments 6–15 mm, anthers 3–7 mm, pollen ca. 50% fertile; style 20–45 mm, stigma surrounded by anthers at anthesis. |
4-merous, zygomorphic, opening near sunset; floral tube 1.5–5 mm; sepals 6–14 mm; petals white, fading pink to red, slightly unequal, elliptic, 7–13 mm, clawed; stamens presented in lower 1/2 of flower, filaments 4.5–11 mm, anthers 2–4.5 mm, pollen 90–100% fertile; style 9–19 mm, stigma exserted beyond anthers at anthesis. |
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Capsules | erect or slightly spreading, usually rusty brown when dry, narrowly lanceoloid to lanceoloid, 15–40 × 4–8 mm, free tips of valves 0.5 mm. |
lanceoloid to narrowly ovoid, 4-winged, 9–19 × 1–3.5 mm, abruptly constricted to a long, sterile stipe 2–10 mm. |
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Seeds | 1.1–1.2 ×0.8–1 mm. |
(1 or)2–4, 2–3(–4) × 0.8–1.3 mm, yellowish to light brown or rarely reddish brown. |
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2n | = 14. |
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Oenothera oakesiana |
Oenothera cinerea |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Sandy coastal meadows and dunes, gravelly or rocky sites along rivers, disturbed sites, roadsides. | |||||
Elevation | 0–50(–500) m. (0–200(–1600) ft.) | |||||
Distribution |
CT; DC; DE; IL; IN; MA; MD; ME; MI; MN; NH; NJ; NY; OH; PA; RI; VA; VT; WI; MB; NB; NF; NS; ON; PE; QC [Introduced in Europe, Asia]
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sc United States
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Discussion | Oenothera oakesiana is a PTH species and forms a ring of 14 chromosomes or a ring of 12 and 1 bivalent in meiosis, and is self-compatible and autogamous (W. Dietrich et al. 1997). It has plastome IV and a AC genome composition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). P. H. Raven and D. P. Gregory (1972[1973]) determined Oenothera cinerea to be self-incompatible. The two subspecies recognized here have disjunct distributions but are very similar morphologically. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Oenothera | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Stipogaura | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | O. biennis var. oakesiana, O. ammophila, O. ammophiloides, O. ammophiloides var. angustifolia, O. ammophiloides var. laurensis, O. atrovirens var. ostreae, O. canovirens var. cymatilis, O. cruciata var. stenopetala, O. cymatilis, O. deflexa var. bracteata, O. disjuncta, O. eriensis, O. eriensis var. niagarensis, O. eriensis var. repandodentata, O. germanica, O. insignis, O. laevigata var. rubripunctata, O. leucophylla, O. litorea, O. magdalena, O. millersii, O. muricata var. parviflora, O. niagarensis, O. nobska, O. oakesiana var. nobska, O. oakesiana var. tidestromii, O. ostreae, O. parviflora var. oakesiana, O. perangusta, O. perangusta var. rubricalyx, O. repandodentata, O. rubescens, O. stenopetala, O. tidestromii, Onagra oakesiana | Gaura cinerea | ||||
Name authority | (A. Gray) J. W. Robbins ex S. Watson & J. M. Coulter in A. Gray et al.: Manual ed. 6, 190. (1890) | (Wooton & Standley) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 211. (2007) | ||||
Web links |