Oenothera mckelveyae |
Oenothera howardii |
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Mckelvey's beeblossom |
Howard's evening-primrose |
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Habit | Herbs perennial, clumped, long-villous, more sparsely so distally, hairs erect, 2–4 mm, also strigillose, sometimes glabrate distally or also sparsely glandular puberulent; from twisted, woody rootstock. | Herbs acaulescent or sometimes caulescent, moderately to densely strigillose and glandular puberulent, sometimes also sparsely to moderately hirsute; from a taproot, sometimes lateral roots producing adventitious shoots. |
Stems | ascending, branched below or just above ground, branched also proximal to inflorescences, 30–70(–120) cm. |
(when present) ascending, longer ones becoming decumbent, leafy, sometimes densely so, 0–10(–30) cm. |
Leaves | in a basal rosette and cauline, basal 3–17 × 0.8–2 cm, blade oblanceolate, cauline 1–6.5 × 0.1–1.5 cm, sessile, blade narrowly oblanceolate to elliptic, margins conspicuously sinuate-dentate, often undulate. |
in a basal rosette, sometimes also cauline, (6–)8.5–17(–23) × (0.5–)1–2(–3) cm; petiole 2–7.5 cm; blade usually oblanceolate, elliptic to narrowly oblong, rarely lanceolate, margins often undulate, entire or remotely and irregularly pinnately lobed mostly in proximal 1/2, rarely more regularly pinnately lobed and lobing extending to distal 1/2, sinuses usually extending less than 1/2 to midrib, lobes triangular to oblong or linear, (1–)4–9(–13) mm, apex acute to obtuse. |
Inflorescences | slender. |
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Flowers | 4-merous, zygomorphic, opening near sunset; floral tube 2–3.5 mm; sepals 6–12 mm; petals white, fading dark pink to red, slightly unequal, elliptic-obovate, 7–11 mm, long-clawed; stamens presented in lower 1/2 of flower, filaments 5–9 mm, lanate at very base, anthers 2–4 mm, pollen 90–100% fertile; style 9–16 mm, stigma exserted beyond anthers at anthesis. |
usually 1 or 2, rarely more, opening per day near sunset, strongly and sweetly scented; buds with unequal free tips 1–3(–4) mm; floral tube (43–)60–110(–125) mm; sepals (30–)35–60(–80) mm; petals brilliant yellow, fading deep red, drying deep reddish purple to reddish brown, usually broadly obovate, rarely subrhombic, (30–)40–60(–73) mm, sometimes with a terminal tooth; filaments (19–)25–38 mm, anthers 10–17 mm; style(90–)110–145(–165) mm, stigma exserted beyond anthers at anthesis. |
Capsules | reflexed, lanceoloid to narrowly ovoid, narrowly 4-winged, 8–19 × 1.5–2 mm, tapering to a sterile stipe 3–9 mm. |
leathery, ovoid, narrowly ovoid, or narrowly lanceoloid to broadly ellipsoid, winged, wings (2–)4–7(–11) mm wide, body (20–)25–50(–80) × 4–6 mm, dehiscent 1/4–1/3 their length; pedicel 2–6 mm. |
Seeds | (1 or)2–4, 2–3 × 1 mm, yellowish to reddish brown. |
numerous, usually in 1 row per locule, rarely in 2 rows toward base, obovoid to subcuboid, 3–8 × 2.5–3.5 mm. |
2n | = 14. |
= 28, 42, 56. |
Oenothera mckelveyae |
Oenothera howardii |
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Phenology | Flowering Mar–Jun. | Flowering May–Jul. |
Habitat | Sandy soil. | Open or rocky areas, in shale, fine-textured sandstones, clays, gypsum, or limestone from High Plains grasslands, open sites in pinyon-juniper woodlands, ponderosa pine-Douglas fir forests. |
Elevation | 0–300 m. (0–1000 ft.) | (1000–)1500–2300(–3000) m. ((3300–)4900–7500(–9800) ft.) |
Distribution |
TX; Mexico (Nuevo León, Tamaulipas) |
CO; KS; NV; UT; WY
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Discussion | Oenothera mckelveyae, on the Rio Grande Plain, is found in an area bounded by from Dimmit and LaSalle counties east to Karnes and Refugio counties in the north, southward through south Texas, extending to northeastern Tamaulipas and adjacent Nuevo León. P. H. Raven and D. P. Gregory (1972[1973]) found Oenothera mckelveyae to be self-incompatible. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenotherahowardii is known from three disjunct areas: three collections on the High Plains (Baca and Otero counties, Colorado, and Hamilton County, Kansas); open yucca-juniper grassland, rocky slopes or disturbed areas on shale substrates along the Colorado counties of Boulder, Denver, Jefferson, and Larimer, and just over the state line in Wyoming; and, common to scattered, mostly on rocky slopes but also in shaded canyon sites on fine-textured red sandstones, clays, gypsum, chalky white degraded limestone or limestone in pinyon-juniper woodland to ponderosa pine-Douglas fir forest in southern Utah and eastern Nevada. A. Nelson intended to publish Lavauxia howardii as a new combination and base it on Oenothera howardii M. E. Jones (1893), which was not validly published at the time, but inadvertently published L. howardii as a new species. Oenothera howardii (A. Nelson) W. L. Wagner is an isonym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Gaura villosa var. mckelveyae, Gaura mckelveyae | Lavauxiahowardii a. |
Name authority | (Munz) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 213. (2007) | (A. Nelson) M. E. Jones ex Prain in B. D. Jackson et al.: Index Kew., suppl. 3: 121. (1908) — (as howardi) |
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