Oenothera macrocarpa |
Oenothera toumeyi |
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bigfruit evening primrose, Missouri evening primrose, Ozark sundrop |
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Habit | Herbs caulescent, strigillose or glabrous, sometimes glandular puberulent distally; from a stout taproot, sometimes lateral roots producing adventitious shoots. | Herbs perennial or sometimes annual, glabrate to strigillose throughout; from a stout taproot. | ||||||||||||||||
Stems | moderately leafy, (1–)4–40(–60) cm. |
1–several, ascending to erect, unbranched to densely branched, 15–70 cm. |
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Leaves | cauline, (2.8–)3.7–12.5(–17) × (0.1–)0.4–3(–4.5) cm; petiole (0.4–)1–4(–6) cm; blade linear, lanceolate-elliptic, elliptic to oblanceolate or suborbiculate, margins entire or conspicuously or inconspicuously denticulate or serrulate, sometimes undulate, apex usually acute, sometimes obtuse or retuse (subsp. incana). |
1–3.5 × 0.1–0.7 cm, fascicles of small leaves 0.2–2.5 cm present in non-flowering axils; petiole 0 cm; blade narrowly lanceolate, base acute-attenuate, margins entire or obscurely and sparsely serrulate, not undulate, apex acute. |
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Flowers | usually 1 or 2, rarely more, opening per day near sunset, fading next morning, sometimes (subspp. |
usually 1 per stem opening per day at sunset; buds with free tips 2–9(–12) mm; floral tube (15–)30–60(–70) mm, funnelform in distal 1/2 or less; sepals 10–25 mm; petals yellow, fading pale pink or pale purple, 10–20 mm; filaments 4–12 mm, anthers 6–10 mm, pollen 85–100% fertile; style 35–70(–80) mm, stigma yellow, quadrangular, usually exserted beyond anthers. |
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Capsules | papery in age, narrowly ellipsoid to lanceoloid, sometimes twisted (subsp. fremontii), winged, wings (2–)10–28(–34) mm wide, body (13–)25–70(–115) × 2–9 mm, dehiscent 1/4–1/3 their length; pedicel 1–12(–25) mm. |
10–50 × 1.5–4 mm, somewhat papery, promptly dehiscent in distal 1/2. |
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Seeds | numerous, rarely as few as 8, in 1 row per locule, obovoid, (2–)3–5 × 1–2.3 mm. |
obovoid, 2–3 mm. |
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Macrocarpa | and oklahomensis) lasting for 2 days, weakly scented; buds with unequal free tips 1–11(–15) mm; floral tube (21–)35–140(–160) mm; sepals (20–)25–65(–75) mm; petals bright yellow, fading orange, reddish orange or mostly unchanged, obovate to very broadly obovate, (17–)25–65(–68) mm, usually with terminal notch and/or tooth, margin sometimes erose; filaments 13–40(–44) mm, anthers 10–24(–25) mm; style (45–)55–192 mm, stigma usually exserted beyond anthers at anthesis. |
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2n | = 14. |
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Oenothera macrocarpa |
Oenothera toumeyi |
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Phenology | Flowering (May–)Jul–Oct. | |||||||||||||||||
Habitat | Local and uncommon on shaded, rocky slopes or disturbed areas, pine-oak forests. | |||||||||||||||||
Elevation | 1500–2600 m. (4900–8500 ft.) | |||||||||||||||||
Distribution |
c United States; n Mexico; s United States
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AZ; NM; Mexico (Chihuahua, Sonora) |
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Discussion | Subspecies 5 (4 in the flora). Oenotheramacrocarpa is variable and has differentiated extensively in the Great Plains region. Each of the five distinctive subspecies occupies a different geographical and ecological situation. Only subsp. mexicana W. L. Wagner from Coahuila, Mexico, occurs outside of the flora area. In general, the subspecies are sharply distinct and each is characterized by a number of features, including pubescence, leaf features, flower and floral tube size, and size and morphology of the capsules and seeds. The five entities are treated as subspecies primarily because of their complete interfertility and extensive intergradation in any area of marginal contact. Intermediates are known between subsp. macrocarpa and subspp. fremontii and oklahomensis and between subspp. incana and oklahomensis. There is also some evidence that suggests past hybridization between subspp. fremontii and incana although there is no present contact between them. All subspecies are self-incompatible. Oenothera alata Nuttall (1818) is an illegitimate name based on O. macrocarpa and pertains here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenothera toumeyi occurs locally from the Chiricahua, Huachuca, and Santa Rita mountains in Cochise and Santa Cruz counties, Arizona, and the Mogollon Mountains in southern Catron County, New Mexico, south through northeastern Sonora in the Sierra Madre Occidental to west-central Chihuahua. H. F. Towner (1977) found that O. toumeyi is self-incompatible and vespertine. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Megapterium | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Calylophus > subsect. Salpingia | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Megapterium macrocarpum, M. nuttallianum | Galpinsia toumeyi, Calylophus hartwegii subsp. toumeyi, C. hartwegii var. toumeyi, C. toumeyi, O. hartwegii var. toumeyi | ||||||||||||||||
Name authority | Nuttall: Cat. Pl. Upper Louisiana, no. 56. (1813) | (Small) Tidestrom: Proc. Biol. Soc. Wash. 48: 41. (1935) | ||||||||||||||||
Web links |