Oenothera macrocarpa subsp. oklahomensis
Onagraceae tribe Onagreae
several, unbranched, sometimes with shorter secondary branches, 5–30(–42) cm.
green, usually flecked with reddish purple splotches or reddish tinged, (5.5–)7–11(–12.5) × (0.3–)0.8–2(–3) cm;
blade usually elliptic to lanceolate-elliptic, sometimes linear, moderately thick, fleshy, margins usually undulate, usually conspicuously denticulate to serrulate, rarely subentire, apex acute.
alternate or basal;
stipules absent.
buds with unequal free tips (5–)6–10(–15) mm;
floral tube (82–)90–140(–147) mm;
sepals 35–60 mm;
petals (38–)42–55(–65) mm;
filaments 24–34 mm, anthers (15–)19–21(–25) mm;
style (120–)140–190 mm.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
globose to narrowly ellipsoid, not twisted, wings (10–)15–20(–25) mm wide, body 35–55(–75) × 7–9 mm.2n = 14.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
Oenothera macrocarpa subsp. oklahomensis
Onagraceae tribe Onagreae
Subspecies oklahomensis is known from Barber, Harper, Meade, and Montgomery counties in extreme southern Kansas south across central Oklahoma to Harmon County in southwestern Oklahoma, and Cooke and Knox counties in adjacent Texas.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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