Oenothera howardii |
Oenothera brachycarpa |
|
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Howard's evening-primrose |
shortfruit evening-primrose |
|
Habit | Herbs acaulescent or sometimes caulescent, moderately to densely strigillose and glandular puberulent, sometimes also sparsely to moderately hirsute; from a taproot, sometimes lateral roots producing adventitious shoots. | Herbs acaulescent or sometimes caulescent, strigillose, also hirsute, hairs often with reddish purple pustulate base, glandular puberulent distally; from a woody taproot, sometimes lateral roots producing adventitious shoots. |
Stems | (when present) ascending, longer ones becoming decumbent, leafy, sometimes densely so, 0–10(–30) cm. |
(when present) ascending, longer ones becoming decumbent, usually densely leafy, 0–20(–36) cm. |
Leaves | in a basal rosette, sometimes also cauline, (6–)8.5–17(–23) × (0.5–)1–2(–3) cm; petiole 2–7.5 cm; blade usually oblanceolate, elliptic to narrowly oblong, rarely lanceolate, margins often undulate, entire or remotely and irregularly pinnately lobed mostly in proximal 1/2, rarely more regularly pinnately lobed and lobing extending to distal 1/2, sinuses usually extending less than 1/2 to midrib, lobes triangular to oblong or linear, (1–)4–9(–13) mm, apex acute to obtuse. |
in a basal rosette, sometimes also cauline, (3.1–)5–21(–34) × (0.3–)1.5–3.5(–5.3) cm; petiole (0.8–)2.5–11(–15) cm; blade usually lanceolate to elliptic, rhombic-obovate, sometimes suborbicular or linear, usually irregularly pinnatifid, some sinuses extending nearly to midrib, usually with a large terminal lobe (0.1–)1.5–2(–2.4) cm, margins erose, apex acute to obtuse or rounded. |
Flowers | usually 1 or 2, rarely more, opening per day near sunset, strongly and sweetly scented; buds with unequal free tips 1–3(–4) mm; floral tube (43–)60–110(–125) mm; sepals (30–)35–60(–80) mm; petals brilliant yellow, fading deep red, drying deep reddish purple to reddish brown, usually broadly obovate, rarely subrhombic, (30–)40–60(–73) mm, sometimes with a terminal tooth; filaments (19–)25–38 mm, anthers 10–17 mm; style(90–)110–145(–165) mm, stigma exserted beyond anthers at anthesis. |
usually 1–3, rarely more, opening per day near sunset, weakly scented; buds with unequal free tips 1–7 mm; floral tube (90–)120–210(–220) mm; sepals 38–55 mm; petals pale yellow to yellow, fading pale orange to pink, drying lavender to purple, usually broadly rhombic-obovate, sometimes obovate, (38–)45–58(–62) mm, distal margin usually erose; filaments (16–)20–32 mm, anthers (8–)13–21 mm; style (123–)155–240(–255) mm, stigma exserted beyond anthers at anthesis. |
Capsules | leathery, ovoid, narrowly ovoid, or narrowly lanceoloid to broadly ellipsoid, winged, wings (2–)4–7(–11) mm wide, body (20–)25–50(–80) × 4–6 mm, dehiscent 1/4–1/3 their length; pedicel 2–6 mm. |
leathery or corky, ovoid to narrowly ellipsoid, ± winged, wings 0–3(–5) mm wide, sometimes capsule with corky thickening between wings, then capsule only 4-angled, body (12–)18–40 × 6–10 mm, dehiscent 1/4 their length; pedicel 0–3 mm. |
Seeds | numerous, usually in 1 row per locule, rarely in 2 rows toward base, obovoid to subcuboid, 3–8 × 2.5–3.5 mm. |
usually numerous, in 1 or 2 rows per locule, obovoid to subcuboid, 3–5 × 1.8–2.2 mm. |
2n | = 28, 42, 56. |
= 14. |
Oenothera howardii |
Oenothera brachycarpa |
|
Phenology | Flowering May–Jul. | Flowering Mar–Aug. |
Habitat | Open or rocky areas, in shale, fine-textured sandstones, clays, gypsum, or limestone from High Plains grasslands, open sites in pinyon-juniper woodlands, ponderosa pine-Douglas fir forests. | Rocky sites, usually on limestone, shale, or gypsum, on igneous substrates from canyons and slopes in Chihuahuan Desert scrub, grasslands, oak-pine-juniper woodlands, open sites in ponderosa pine-Douglas fir forests. |
Elevation | (1000–)1500–2300(–3000) m. ((3300–)4900–7500(–9800) ft.) | 1000–2700 m. (3300–8900 ft.) |
Distribution |
CO; KS; NV; UT; WY
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AZ; NM; TX; Mexico (Chihuahua, Coahuila, Durango, Sonora)
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Discussion | Oenotherahowardii is known from three disjunct areas: three collections on the High Plains (Baca and Otero counties, Colorado, and Hamilton County, Kansas); open yucca-juniper grassland, rocky slopes or disturbed areas on shale substrates along the Colorado counties of Boulder, Denver, Jefferson, and Larimer, and just over the state line in Wyoming; and, common to scattered, mostly on rocky slopes but also in shaded canyon sites on fine-textured red sandstones, clays, gypsum, chalky white degraded limestone or limestone in pinyon-juniper woodland to ponderosa pine-Douglas fir forest in southern Utah and eastern Nevada. A. Nelson intended to publish Lavauxia howardii as a new combination and base it on Oenothera howardii M. E. Jones (1893), which was not validly published at the time, but inadvertently published L. howardii as a new species. Oenothera howardii (A. Nelson) W. L. Wagner is an isonym. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Oenotherabrachycarpa occurs from southeastern Arizona in southern Navajo, southeastern Pima, Graham, Santa Cruz, and Cochise counties, east across southern New Mexico to Val Verde and Pecos counties in trans-Pecos Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Megapterium | Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Megapterium |
Sibling taxa | ||
Synonyms | Lavauxiahowardii a. | Lavauxia brachycarpa, L. graminifolia, L. wrightii, Megapterium brachycarpum, M. brachycarpum var. wrightii, O. australis, O. brachycarpa var. wrightii, O. cespitosa subsp. australis, O. cespitosa var. australis, O. graminifolia, O. wrightii, Pachylophus australis |
Name authority | (A. Nelson) M. E. Jones ex Prain in B. D. Jackson et al.: Index Kew., suppl. 3: 121. (1908) — (as howardi) | A. Gray: Smithsonian Contr. Knowl. 3(5): 70. (1852) |
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