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variableleaf evening primrose

demaree's beeblossom

Habit Herbs annual or short-lived perennial, sparsely to densely strigillose, inflorescence sometimes also sparsely glandular puberulent, villous, or sparsely hirsute with spreading, pustulate-based hairs, or sometimes glabrate. Herbs usually robust winter-annual, sometimes biennial, densely strigillose throughout; from fleshy taproot.
Stems

unbranched or branched mainly in distal part, 25–70 cm.

usually well-branched distal to base, 50–400 cm.

Leaves

in a basal rosette and cauline, basal 7–15 × 1–2.5 cm, cauline 3–13 ×0.4–2.3 cm;

blade narrowly oblanceolate to oblanceolate, gradually narrowly lanceolate to lanceolate or elliptic distally, margins deeply lobed to remotely dentate or subentire;

bracts longer than capsule they subtend, 1–3 cm.

in a basal rosette and cauline, 3–7 × 0.2–0.8 cm;

blade narrowly elliptic to elliptic or lanceolate, margins subentire or shallowly undulate-denticulate.

Inflorescences

dense, often with several lateral branches, mature buds usually overtopping spike apex.

Flowers

2–several per spike opening per day near sunset;

buds erect, with free tips erect or spreading, 1–6 mm;

floral tube nearly straight, 25–47 mm;

sepals 15–30 mm;

petals yellow, broadly elliptic to nearly rhombic, 18–35 mm;

filaments 15–30 mm, anthers 3–8 mm, pollen 85–100% fertile;

style 45–75 mm, stigma usually exserted beyond anthers at anthesis.

4-merous, zygomorphic, opening at sunrise;

floral tube 4–13(–15) mm;

sepals 13–20 mm;

petals white, fading pink, rhombic-obovate, 10–17 mm;

filaments 8–17 mm, anthers 3–7 mm, pollen 90–100% fertile;

style 18–32 mm, stigma exserted beyond anthers at anthesis.

Capsules

lanceoloid, 13–25 × 2.5–4 mm.

ellipsoid or ovoid, sharply 4-angled, 4.5–7 × 1.5–2.5 mm;

sessile.

Seeds

brown, often flecked with darker spots, ellipsoid to broadly ellipsoid, 1.1–1.8 × 0.4–0.8 mm.

2–4, yellowish to reddish brown, 1.2–3 × 0.7–1.3 mm.

2n

= 14.

Oenothera heterophylla

Oenothera demareei

Phenology Flowering Jul–Oct.
Habitat Open meadows in sandy loam.
Elevation 70–200 m. (200–700 ft.)
Distribution
from USDA
s United States
[BONAP county map]
from FNA
AR
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Oenothera demareei is known only from Clark, Garland, Hempstead, Howard, Montgomery, Pike, Saline, and Sevier counties.

P. H. Raven and D. P. Gregory (1972[1973]) found Oenothera demareei to be self-incompatible.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Herbs densely to sparsely strigillose, also at least parts of inflorescence sparsely hirsute with spreading, pustulate-based hairs, and often glandular puberulent and villous; flower buds with free tips spreading, 2–6 mm.
subsp. heterophylla
1. Herbs densely to sparsely strigillose, also at least part of inflorescence glabrate or glandular puberulent; flower buds with free tips erect, 1–3 mm.
subsp. orientalis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Oenothera > subsect. Candela Onagraceae > subfam. Onagroideae > tribe Onagreae > Oenothera > sect. Gaura > subsect. Gaura
Sibling taxa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. demareei, O. dodgeniana, O. drummondii, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glaucifolia, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. havardii, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. nealleyi, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
O. acutissima, O. albicaulis, O. argillicola, O. arida, O. arizonica, O. biennis, O. boquillensis, O. brachycarpa, O. calcicola, O. californica, O. canescens, O. capillifolia, O. cavernae, O. cespitosa, O. cinerea, O. clelandii, O. coloradensis, O. cordata, O. coronopifolia, O. coryi, O. curtiflora, O. curtissii, O. deltoides, O. dodgeniana, O. drummondii, O. elata, O. engelmannii, O. falfurriae, O. filiformis, O. filipes, O. flava, O. fruticosa, O. gaura, O. gayleana, O. glaucifolia, O. glazioviana, O. grandiflora, O. grandis, O. harringtonii, O. hartwegii, O. havardii, O. heterophylla, O. hispida, O. howardii, O. humifusa, O. jamesii, O. kunthiana, O. laciniata, O. lavandulifolia, O. lindheimeri, O. linifolia, O. longissima, O. macrocarpa, O. mckelveyae, O. mexicana, O. nealleyi, O. neomexicana, O. nutans, O. nuttallii, O. oakesiana, O. organensis, O. pallida, O. parviflora, O. patriciae, O. perennis, O. pilosella, O. platanorum, O. podocarpa, O. primiveris, O. psammophila, O. pubescens, O. rhombipetala, O. riparia, O. rosea, O. serrulata, O. sessilis, O. simulans, O. sinuosa, O. spachiana, O. speciosa, O. stricta, O. suffrutescens, O. suffulta, O. tetraptera, O. texensis, O. toumeyi, O. triangulata, O. triloba, O. tubicula, O. villosa, O. wolfii, O. xylocarpa
Subordinate taxa
O. heterophylla subsp. heterophylla, O. heterophylla subsp. orientalis
Synonyms Raimannia heterophylla Gaura demareei
Name authority Spach: Nouv. Ann. Mus. Hist. Nat. 4: 348. (1836) (P. H. Raven & D. P. Gregory) W. L. Wagner & Hoch: Syst. Bot. Monogr. 83: 212. (2007)
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